第19章
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  Raphanusraphanistrumandsativushaveoftenbeenrankedasdistinctspecies,andowingtodifferencesintheirfruitevenasdistinctgenera;

  butProfessorHoffman(’Bot。Zeitung’1872page482)hasnowshownthatthesedifferences,remarkableastheyare,graduateaway,thefruitofR。

  caudatusbeingintermediate。BycultivatingR。raphanistrumduringseveralgenerations(ibid1873page9),ProfessorHoffmanalsoobtainedplantsbearingfruitslikethoseofR。sativus。

  PEA(Pisumsativum)。

  Mostbotanistslookatthegarden—peaasspecificallydistinctfromthefield—pea(P。arvense)。ThelatterexistsinawildstateinSouthernEurope;buttheaboriginalparentofthegarden—peahasbeenfoundbyonecollectoralone,ashestates,intheCrimea。(9/81。Alph。DeCandolle’Geograph。Bot。’960。Mr。Bentham’Hort。Journal’volume91855page141

  believesthatgardenandfieldpeasbelongtothesamespecies,andinthisrespecthediffersfromDr。Targioni。)AndrewKnightcrossed,asIaminformedbytheRev。A。Fitch,thefield—peawithawell—knowngardenvariety,thePrussianpea,andthecrossseemstohavebeenperfectlyfertile。Dr。Alefieldhasrecentlystudied(9/82。’BotanischeZeitung’1860

  s。204。)thegenuswithcare,and,afterhavingcultivatedaboutfiftyvarieties,concludesthatcertainlytheyallbelongtothesamespecies。Itisaninterestingfactalreadyalludedto,that,accordingtoO。Heer(9/83。’DiePflanzenderPfahlbauten’1866s。23。)thepeasfoundinthelake—habitationsofSwitzerlandoftheStoneandBronzeages,belongtoanextinctvariety,withexceedinglysmallseeds,alliedtoP。arvenseorthefield—pea。Thevarietiesofthecommongarden—peaarenumerous,anddifferconsiderablyfromoneanother。ForcomparisonIplantedatthesametimeforty—one,EnglishandFrenchvarieties。Theydifferedgreatlyinheight,——

  namelyfrombetween6and12inchesto8feet(9/84。AvarietycalledtheRouncivaattainsthisheight,asisstatedbyMr。Gordonin’Transact。

  Hort。Soc。’2ndseriesvolume11835page374fromwhichpaperIhavetakensomefacts。),——inmannerofgrowth,andinperiodofmaturity。Somedifferingeneralaspectevenwhileonlytwoorthreeinchesinheight。ThestemsofthePrussianpeaaremuchbranched。Thetallkindshavelargerleavesthanthedwarfkinds,butnotinstrictproportiontotheirheight:——HAIR’S

  DWARFMONMOUTHhasverylargeleaves,andthePOISNAINHATIF,andthemoderatelytallBLUEPRUSSIAN,haveleavesabouttwo—thirdsofthesizeofthetallestkind。IntheDANECROFTtheleafletsarerathersmallandalittlepointed;intheQUEENOFDWARFSratherrounded;andintheQUEENOF

  ENGLANDbroadandlarge。Inthesethreepeastheslightdifferencesintheshapeoftheleavesareaccompaniedbyslightdifferencesincolour,inthePOISGEANTSANSPARCHEMIN,whichbearspurpleflowers,theleafletsintheyoungplantareedgedwithred;andinallthepeaswithpurpleflowersthestipulesaremarkedwithred。

  Inthedifferentvarieties,one,two,orseveralflowersinasmallcluster,areborneonthesamepeduncle;andthisisadifferencewhichisconsideredofspecificvalueinsomeoftheLeguminosae。Inallthevarietiestheflowerscloselyresembleeachotherexceptincolourandsize。Theyaregenerallywhite,sometimespurple,butthecolourisinconstanteveninthesamevariety。InWARNER’SEMPEROR,whichisatallkind,theflowersarenearlydoublethesizeofthePOISNAINHATIF;butHAIR’SDWARFMONMOUTH,whichhaslargeleaves,likewisehaslargeflowers。

  ThecalyxintheVICTORIAMARROWislarge,andinBISHOP’SLONGPODthesepalsarerathernarrow。Innootherkindisthereanydifferenceintheflower。

  Thepodsandseeds,whichwithnaturalspeciesaffordsuchconstantcharacters,differgreatlyinthecultivatedvarietiesofthepea;andthesearethevaluable,andconsequentlytheselectedparts。SUGARPEAS,orPOISSANSPARCHEMIN,areremarkablefromtheirthinpods,which,whilstyoung,arecookedandeatenwhole;andinthisgroup,which,accordingtoMr。Gordonincludeselevensub—varieties,itisthepodwhichdiffersmost;

  thusLEWIS’SNEGRO—PODDEDPEAhasastraight,broad,smooth,anddark—

  purplepod,withthehusknotsothinasintheotherkinds;thepodofanothervarietyisextremelybowed;thatofthePOISGEANTismuchpointedattheextremity;andinthevariety\"AGRANDSCOSSES\"thepeasareseenthroughthehuskinsoconspicuousamannerthatthepod,especiallywhendry,canhardlyatfirstberecognisedasthatofapea。

  Intheordinaryvarietiesthepodsalsodiffermuchinsize;——incolour,thatofWOODFORD’SGREENMARROWbeingbright—greenwhendry,insteadofpalebrown,andthatofthepurple—poddedpeabeingexpressedbyitsname;—

  —insmoothness,thatofDANECROFTbeingremarkablyglossy,whereasthatoftheNEPLUSULTRAisrugged;inbeingeithernearlycylindrical,orbroadandflat;——inbeingpointedattheend,asinTHURSTON’SRELIANCE,ormuchtruncated,asintheAMERICANDWARF。IntheAUVERGNEPEAthewholeendofthepodisbowedupwards。IntheQUEENOFTHEDWARFSandinSCIMITARPEAS

  thepodisalmostellipticinshape。Iheregivedrawingsofthefourmostdistinctpodsproducedbytheplantscultivatedbyme。

  (FIGURE41。PODSANDPEAS。I。QueenofDwarfs。II。AmericanDwarf。III。

  Thurston’sReliance。IV。PoisGeantsansparchemin。a。DanO’RourkePea。b。

  QueenofDwarfsPea。c。Knight’sTallWhiteMarrow。a。Lewis’sNegroPea。)

  Inthepeaitselfwehaveeverytintbetweenalmostpurewhite,brown,yellow,andintensegreen;inthevarietiesoftheSUGARPEASwehavethesesametints,togetherwithredpassingthroughfinepurpleintoadarkchocolatetint。Thesecoloursareeitheruniformordistributedindots,striae,ormoss—likemarks;theydependinsomecasesonthecolourofthecotyledonsseenthroughtheskin,andinothercasesontheoutercoatsofthepeaitself。Inthedifferentvarieties,thepodscontain,accordingtoMr。Gordon,fromelevenortwelvetoonlyfourorfivepeas。Thelargestpeasarenearlytwiceasmuchindiameterasthesmallest;andthelatterarenotalwaysbornebythemostdwarfedkinds。Peasdiffermuchinshape,beingsmoothandspherical,smoothandoblong,nearlyovalintheQUEENOF

  THEDWARFS,andnearlycubicalandcrumpledinmanyofthelargerkinds。

  Withrespecttothevalueofthedifferencesbetweenthechiefvarieties,itcannotbedoubtedthat,ifoneofthetallSUGAR—PEAS,withpurpleflowers,thin—skinnedpodsofanextraordinaryshape,includinglarge,dark—purplepeas,grewwildbythesideofthelowlyQUEENOFTHEDWARFS,withwhiteflowers,greyish—green,roundedleaves,scimitar—likepods,containingoblong,smooth,pale—colouredpeas,whichbecamematureatadifferentseason:orbythesideofoneofthegiganticsorts,liketheCHAMPIONOFENGLAND,withleavesofgreatsize,pointedpods,andlarge,green,crumpled,almostcubicalpeas,——allthreekindswouldberankedasdistinctspecies。

  AndrewKnight(9/85。’Phil。Tract。’1799page196。)hasobservedthatthevarietiesofpeaskeepverytrue,becausetheyarenotcrossedbyinsects。

  Asfarasthefactofkeepingtrueisconcerned,IhearfromMr。MastersofCanterbury,wellknownastheoriginatorofseveralnewkinds,thatcertainvarietieshaveremainedconstantforaconsiderabletime,——forinstance,KNIGHT’SBLUEDWARF,whichcameoutabouttheyear1820。(9/86。’Gardener’sMagazine’volume11826page153。)Butthegreaternumberofvarietieshaveasingularlyshortexistence:thusLoudonremarks(9/87。’EncyclopaediaofGardening’page823。)that\"sortswhichwerehighlyapprovedin1821,arenow,in1833,nowheretobefound;\"andoncomparingthelistsof1833withthoseof1855,Ifindthatnearlyallthevarietieshavechanged。Mr。

  Mastersinformsmethatthenatureofthesoilcausessomevarietiestolosetheircharacter。Aswithotherplants,certainvarietiescanbepropagatedtruly,whilstothersshowadeterminedtendencytovary;thustwopeasdifferinginshape,oneroundandtheotherwrinkled,werefoundbyMr。Masterswithinthesamepod,buttheplantsraisedfromthewrinkledkindalwaysevincedastrongtendencytoproduceroundpeas。Mr。Mastersalsoraisedfromaplantofanothervarietyfourdistinctsub—varieties,whichboreblueandround,whiteandround,blueandwrinkled,andwhiteandwrinkledpeas;andalthoughhesowedthesefourvarietiesseparatelyduringseveralsuccessiveyears,eachkindalwaysreproducedallfourkindsmixedtogether!

  Withrespecttothevarietiesnotnaturallyintercrossing,Ihaveascertainedthatthepea,whichinthisrespectdiffersfromsomeotherLeguminosae,isperfectlyfertilewithouttheaidofinsects。YetIhaveseenhumble—beeswhilstsuckingthenectardepressthekeel—petals,andbecomesothicklydustedwithpollen,thatitcouldhardlyfailtobeleftonthestigmaofthenextflowerwhichwasvisited。Nevertheless,distinctvarietiesgrowingcloselytogetherrarelycross;andIhavereasontobelievethatthisisduetotheirstigmasbeingprematurelyfertilisedinthiscountrybypollenfromthesameflower。Thehorticulturistswhoraiseseed—peasarethusenabledtoplantdistinctvarietiesclosetogetherwithoutanybadconsequences;anditiscertain,asIhavemyselffound,thattrueseedmaybesavedduringatleastseveralgenerationsunderthesecircumstances。(9/88。SeeDr。Andersontothesameeffectinthe’BathSoc。

  AgriculturalPapers’volume4page87。)Mr。Fitchraised,asheinformsme,onevarietyfortwentyyears,anditalwayscametrue,thoughgrownclosetoothervarieties。Fromtheanalogyofkidney—beansIshouldhaveexpected(9/89。Ihavepublishedfulldetailsofexperimentsonthissubjectinthe’Gardener’sChronicle’1857October25。)thatvarietiesthuscircumstancedwouldhaveoccasionallycrossed;andIshallgiveintheeleventhchaptertwocasesofthishavingoccurred,asshown(inamannerhereaftertobeexplained)bythepollenoftheonevarietyhavingacteddirectlyontheseedsoftheother。Whethermanyofthenewvarietieswhichincessantlyappearareduetosuchoccasionalandaccidentalcrosses,Idonotknow。

  NordoIknowwhethertheshortexistenceofalmostallthenumerousvarietiesistheresultofmerechangeoffashion,oroftheirhavingaweakconstitution,frombeingtheproductoflong—continuedself—

  fertilisation。Itmay,however,benoticedthatseveralofAndrewKnight’svarieties,whichhaveenduredlongerthanmostkinds,wereraisedtowardsthecloseofthelastcenturybyartificialcrosses;someofthem,I

  believe,werestillvigorousin1860;butnow,in1865,awriter,speaking(9/90。’Gardener’sChronicle’1865page387。)ofKnight’sfourkindsofmarrows,says,theyhaveacquiredafamoushistory,buttheirgloryhasdeparted。

  WithrespecttoBeans(Fabavulgaris),Iwillsaybutlittle。Dr。Alefieldhasgiven(9/91。’Bonplandia’10,1862s。348。)shortdiagnosticcharactersoffortyvarieties。Everyonewhohasseenacollectionmusthavebeenstruckwiththegreatdifferenceinshape,thickness,proportionallengthandbreadth,colour,andsizewhichbeanspresent。WhatacontrastbetweenaWindsorandHorse—bean!Asinthecaseofthepea,ourexistingvarietieswereprecededduringtheBronzeageinSwitzerland(9/92。Heer’DiePflanzenderPfahlbauten’1866s。22。)byapeculiarandnowextinctvarietyproducingverysmallbeans。(9/93。Mr。BenthaminformsmethatinPoitouandtheadjoiningpartsofFrance,varietiesofPhaseolusvulgarisareextremelynumerous,andsodifferentthattheyweredescribedbySaviasdistinctspecies。Mr。Benthambelievesthatallaredescendedfromanunknowneasternspecies。Althoughthevarietiesdiffersogreatlyinstatureandintheirseeds,\"thereisaremarkablesamenessintheneglectedcharactersoffoliageandflowers,andespeciallyinthebracteoles,aninsignificantcharacterintheeyesevenofbotanists。\")

  POTATO(Solanumtuberosum)。

  Thereislittledoubtabouttheparentageofthisplant;forthecultivatedvarietiesdifferextremelylittleingeneralappearancefromthewildspecies,whichcanberecognisedinitsnativelandatthefirstglance。

  (9/94。Darwin’JournalofResearches’1845page285。Sabinein’Transact。

  Hort。Soc。’volume5page249。)ThevarietiescultivatedinBritainarenumerous;thusLawson(9/95。’SynopsisoftheVegetableProductsofScotland’quotedinWilson’s’BritishFarming’page317。)givesadescriptionof175kinds。Iplantedeighteenkindsinadjoiningrows;theirstemsandleavesdifferedbutlittle,andinseveralcasestherewasasgreatadifferencebetweentheindividualsofthesamevarietyasbetweenthedifferentvarieties。Theflowervariedinsize,andincolourbetweenwhiteandpurple,butinnootherrespect,exceptthatinonekindthesepalsweresomewhatelongated。Onestrangevarietyhasbeendescribedwhichalwaysproducestwosortsofflowers,thefirstdoubleandsterile,thesecondsingleandfertile。(9/96。SirG。Mackenzie,in’Gardener’sChronicle’1845page790。)Thefruitorberriesalsodiffer,butonlyinaslightdegree。(9/97。PutscheundVertuch’VersucheinerMonographiederKartoffeln’1819s。9,15。SeealsoDr。Anderson’RecreationsinAgriculture’volume4page325。)ThevarietiesareliableinverydifferentdegreetotheattackoftheColoradopotato—beetle。(9/98。Walsh’TheAmericanEntomologist’1869page160。AlsoS。Tenney’TheAmericanNaturalist’May1871page171。)

  Thetubers,ontheotherhand,presentawonderfulamountofdiversity。

  Thisfactaccordswiththeprinciplethatthevaluableandselectedpartsofallcultivatedproductionspresentthegreatestamountofmodification。

  Theydiffermuchinsizeandshape,beingglobular,oval,flattened,kidney—like,orcylindrical。OnevarietyfromPeruisdescribed(9/99。

  ’Gardener’sChronicle’1862page1052。)asbeingquitestraight,andatleastsixinchesinlength,thoughnothickerthanaman’sfinger。Theeyesorbudsdifferinform,position,andcolour。Themannerinwhichthetubersarearrangedontheso—calledrootsorrhizomesisdifferent;thus,inthegurken—kartoffelntheyformapyramidwiththeapexdownwards,andinanothervarietytheyburythemselvesdeepintheground。Therootsthemselvesruneithernearthesurfaceordeepintheground。Thetubersalsodifferinsmoothnessandcolour,beingexternallywhite,red,purple,oralmostblack,andinternallywhite,yellow,oralmostblack。Theydifferinflavourandquality,beingeitherwaxyormealy;intheirperiodofmaturity,andintheircapacityforlongpreservation。

  Aswithmanyotherplantswhichhavebeenlongpropagatedbybulbs,tubers,cuttings,etc。,bywhichmeansthesameindividualisexposedduringalengthoftimetodiversifiedconditions,seedlingpotatoesgenerallydisplayinnumerableslightdifferences。Severalvarieties,evenwhenpropagatedbytubers,arefarfromconstant,aswillbeseeninthechapteronBud—variation。Dr。Anderson(9/100。’BathSocietyAgricult。Papers’

  volume5page127。And’RecreationsinAgriculture’volume5page86。)

  procuredseedfromanIrishpurplepotato,whichgrewfarfromanyotherkind,sothatitcouldnotatleastinthisgenerationhavebeencrossed,yetthemanyseedlingsvariedinalmosteverypossiblerespect,sothat\"scarcelytwoplantswereexactlyalike。\"Someoftheplantswhichcloselyresembledeachotheraboveground,producedextremelydissimilartubers;

  andsometuberswhichexternallycouldhardlybedistinguished,differedwidelyinqualitywhencooked。Eveninthiscaseofextremevariability,theparent—stockhadsomeinfluenceontheprogeny,forthegreaternumberoftheseedlingsresembledinsomedegreetheparentIrishpotato。Kidneypotatoesmustberankedamongstthemosthighlycultivatedandartificialraces;neverthelesstheirpeculiaritiescanoftenbestrictlypropagatedbyseed。Agreatauthority,Mr。Rivers(9/101。’Gardener’sChronicle’1863

  page643。)statesthat\"seedlingsfromtheash—leavedkidneyalwaysbearastrongresemblancetotheirparent。Seedlingsfromthefluke—kidneyarestillmoreremarkablefortheiradherencetotheirparentstock,for,oncloselyobservingagreatnumberduringtwoseasons,Ihavenotbeenabletoobservetheleastdifference,eitherinearliness,productiveness,orinthesizeorshapeoftheirtubers。\"

  CHAPTER1。X。

  PLANTScontinued——FRUITS——ORNAMENTALTREES——FLOWERS。

  FRUITS。

  GRAPES。

  VARYINODDANDTRIFLINGPARTICULARS。

  MULBERRY。

  THEORANGEGROUP。

  SINGULARRESULTSFROMCROSSING。

  PEACHANDNECTARINE。

  BUD—VARIATION。

  ANALOGOUSVARIATION。

  RELATIONTOTHEALMOND。

  APRICOT。

  PLUMS。

  VARIATIONINTHEIRSTONES。

  CHERRIES。

  SINGULARVARIETIESOF。

  APPLE。

  PEAR。

  STRAWBERRY。

  INTERBLENDINGOFTHEORIGINALFORMS。

  GOOSEBERRY。

  STEADYINCREASEINSIZEOFTHEFRUIT。

  VARIETIESOF。

  WALNUT。

  NUT。

  CUCURBITACEOUSPLANTS。

  WONDERFULVARIATIONOF。

  ORNAMENTALTREES。

  THEIRVARIATIONINDEGREEANDKIND。

  ASH—TREE。

  SCOTCH—FIR。

  HAWTHORN。

  FLOWERS。

  MULTIPLEORIGINOFMANYKINDS。

  VARIATIONINCONSTITUTIONALPECULIARITIES。

  KINDOFVARIATION。

  ROSES。

  SEVERALSPECIESCULTIVATED。

  PANSY。

  DAHLIA。

  HYACINTH。

  HISTORYANDVARIATIONOF。

  [THEVINE(Vitisvinifera)(Grapes)。

  ThebestauthoritiesconsiderallourgrapesasthedescendantsofonespecieswhichnowgrowswildinwesternAsia,whichgrewwildduringtheBronzeageinItaly(10/1。Heer’PflanzenderPfahlbauten’1866s。28。),andwhichhasrecentlybeenfoundfossilinatufaceousdepositinthesouthofFrance。(10/2。Alph。DeCandolle’Geograph。Bot。’page872;Dr。A。

  Targioni—Tozzettiin’Jour。Hort。Soc。’vol9page133。ForthefossilvinefoundbyDr。G。Planchonsee’Nat。Hist。Review’1865Aprilpage224。SeealsothevaluableworksofM。deSaportaonthe’TertiaryPlantsofFrance。’)Someauthors,however,entertainmuchdoubtaboutthesingleparentageofourcultivatedvarieties,owingtothenumberofsemi—wildformsfoundinSouthernEurope,especiallyasdescribedbyClemente(10/3。

  Godron’Del’Espece’tome2page100。)inaforestinSpain;butasthegrapesowsitselffreelyinSouthernEurope,andasseveralofthechiefkindstransmittheircharactersbyseed(10/4。SeeanaccountofM。

  Vibert’sexperimentsbyAlex。Jordanin’Mem。del’Acad。deLyon’tome2

  1852page108。),whilstothersareextremelyvariable,theexistenceofmanydifferentescapedformscouldhardlyfailtooccurincountrieswherethisplanthasbeencultivatedfromtheremotestantiquity。Thatthevinevariesmuchwhenpropagatedbyseed,wemayinferfromthelargelyincreasednumberofvarietiessincetheearlierhistoricalrecords。Newhot—housevarietiesareproducedalmosteveryyear;forinstance(10/5。

  ’Gardener’sChronicle’1864page488。)agolden—colouredvarietyhasbeenrecentlyraisedinEnglandfromablackgrapewithouttheaidofacross。

  VanMons(10/6。’ArbresFruitiers’1836tome2page290。)rearedamultitudeofvarietiesfromtheseedofonevine,whichwascompletelyseparatedfromallothers,sothattherecouldnot,atleastinthisgeneration,havebeenanycrossing,andtheseedlingspresented\"lesanaloguesdetouteslessortes,\"anddifferedinalmosteverypossiblecharacterbothinthefruitsandfoliage。

  Thecultivatedvarietiesareextremelynumerous;CountOdartsaysthathewillnotdenythattheremayexistthroughouttheworld700or800,perhapseven1000varieties,butnotathirdofthesehaveanyvalue。InthecatalogueoffruitcultivatedintheHorticulturalGardensofLondon,publishedin1842,99varietiesareenumerated。Whereverthegrapeisgrownmanyvarietiesoccur:Pallasdescribes24intheCrimea,andBurnesmentions10inCabool。Theclassificationofthevarietieshasmuchperplexedwriters,andCountOdartisreducedtoageographicalsystem;butIwillnotenteronthissubject,noronthemanyandgreatdifferencesbetweenthevarieties。Iwillmerelyspecifyafewcuriousandtriflingpeculiarities,alltakenfromOdart’shighlyesteemedwork(10/7。Odart’AmpelographieUniverselle’1849。)forthesakeofshowingthediversifiedvariabilityofthisplant。Simonhasclassedgrapesintotwomaindivisions,thosewithdownyleaves,andthosewithsmoothleaves,butheadmitsthatinonevariety,namelytheRebazo,theleavesareeithersmooth,ordowny;andOdart(page70)statesthatsomevarietieshavethenervesalone,andothervarietiestheiryoungleaves,downy,whilsttheoldonesaresmooth。ThePedro—Ximenesgrape(Odartpage397)presentsapeculiaritybywhichitcanbeatoncerecognisedamongstahostofothervarieties,namely,thatwhenthefruitisnearlyripethenervesoftheleavesoreventhewholesurfacebecomesyellow。TheBarberad’Astiiswellmarkedbyseveralcharacters(page426),amongstothers,\"bysomeoftheleaves,anditisalwaysthelowestonthebranches,suddenlybecomingofadarkredcolour。\"Severalauthorsinclassifyinggrapeshavefoundedtheirmaindivisionsontheberriesbeingeitherroundoroblong;andOdartadmitsthevalueofthischaracter;yetthereisonevariety,theMaccabeo(page71),whichoftenproducessmallround,andlargeoblong,berriesinthesamebunch。CertaingrapescalledNebbiolo(page429)presentaconstantcharacter,sufficientfortheirrecognition,namely,\"theslightadherenceofthatpartofthepulpwhichsurroundstheseedstotherestoftheberry,whencutthroughtransversely。\"ARhenishvarietyismentioned(page228)whichlikesadrysoil;thefruitripenswell,butatthemomentofmaturity,ifmuchrainfalls,theberriesareapttorot;ontheotherhand,thefruitofaSwissvariety(page243)isvaluedforwellsustainingprolongedhumidity。Thislattervarietysproutslateinthespring,yetmaturesitsfruitearly;othervarieties(page362)havethefaultofbeingtoomuchexcitedbytheAprilsun,andinconsequencesufferfromfrost。A

  Styrianvariety(page254)hasbrittlefoot—stalks,sothattheclustersoffruitareoftenblownoff;thisvarietyissaidtobeparticularlyattractivetowaspsandbees。Othervarietieshavetoughstalks,whichresistthewind。Manyothervariablecharacterscouldbegiven,buttheforegoingfactsaresufficienttoshowinhowmanysmallstructuralandconstitutionaldetailsthevinevaries。DuringthevinediseaseinFrancecertainoldgroupsofvarieties(10/8。M。Bouchardatin’ComptesRendus’

  December1,1851quotedin’Gardener’sChronicle’1852page435。SeealsoC。V。RileyonthemannerinwhichsomefewofthevarietiesoftheAmericanLabruscanVineescapetheattacksofthePhylloxera:’FourthAnnualReportontheInsectsofMissouri’1872page63and’FifthReport’1873page66。)

  havesufferedfarmorefrommildewthanothers。Thus\"thegroupofChasselas,sorichinvarieties,didnotaffordasinglefortunateexception;\"certainothergroupssufferedmuchless;thetrueoldBurgundy,forinstance,wascomparativelyfreefromdisease,andtheCarminatlikewiseresistedtheattack。TheAmericanvines,whichbelongtoadistinctspecies,entirelyescapedthediseaseinFrance;andwethusseethatthoseEuropeanvarietieswhichbestresistthediseasemusthaveacquiredinaslightdegreethesameconstitutionalpeculiaritiesastheAmericanspecies。

  WHITEMULBERRY(Morusalba)。

  Imentionthisplantbecauseithasvariedincertaincharacters,namely,inthetextureandqualityoftheleaves,fittingthemtoserveasfoodforthedomesticatedsilkworm,inamannernotobservedwithotherplants;butthishasarisensimplyfromsuchvariationsinthemulberryhavingbeenattendedto,selected,andrenderedmoreorlessconstant。M。deQuatrefages(10/9。’EtudessurlesMaladiesactuellesduVeraSoie’1859

  page321。)brieflydescribessixkindscultivatedinonevalleyinFrance:

  ofthesetheAMOUROUSOproducesexcellentleaves,butisrapidlybeingabandonedbecauseitproducesmuchfruitmingledwiththeleaves:theANTOFINOyieldsdeeplycutleavesofthefinestquality,butnotingreatquantity:theCLAROismuchsoughtforbecausetheleavescanbeeasilycollected:lastly,theROSObearsstronghardyleaves,producedinlargequantity,butwiththeoneinconvenience,thattheyarebestadaptedforthewormsaftertheirfourthmoult。MM。Jacquemet—Bonnefont,ofLyon,however,remarkintheircatalogue(1862)thattwosub—varietieshavebeenconfoundedunderthenameoftheroso,onehavingleavestoothickforthecaterpillars,theotherbeingvaluablebecausetheleavescaneasilybegatheredfromthebrancheswithoutthebarkbeingtorn。

  InIndiathemulberryhasalsogivenrisetomanyvarieties。TheIndianformisthoughtbymanybotaniststobeadistinctspecies;butasRoyleremarks(10/10。’ProductiveResourcesofIndia’page130。),\"somanyvarietieshavebeenproducedbycultivationthatitisdifficulttoascertainwhethertheyallbelongtoonespecies;\"theyare,asheadds,nearlyasnumerousasthoseofthesilkworm。

  THEORANGEGROUP。

  Weheremeetwithgreatconfusioninthespecificdistinctionandparentageoftheseveralkinds。Gallesio(10/11。’TraiteduCitrus’1811。’TeoriadellaRiproduzioneVegetale’1816。Iquotechieflyfromthissecondwork。

  In1839Gallesiopublishedinfolio’GliAgrumideiGiard。Bot。diFirenze’

  inwhichhegivesacuriousdiagramofthesupposedrelationshipofalltheforms。),whoalmostdevotedhislife—timetothesubject,considersthattherearefourspecies,namely,sweetandbitteroranges,lemons,andcitrons,eachofwhichhasgivenrisetowholegroupsofvarieties,monsters,andsupposedhybrids。Onehighauthority(10/12。Mr。Bentham’ReviewofDr。A。Targioni—Tozzetti,JournalofHort。Soc。’volume9page133。)believesthatthesefourreputedspeciesareallvarietiesofthewildCitrusmedica,butthattheshaddock(Citrusdecumana),whichisnotknowninawildstate,isadistinctspecies;thoughitsdistinctnessisdoubtedbyanotherwriter\"ofgreatauthorityonsuchmatters,\"namely,Dr。

  BuchananHamilton。Alph。DeCandolle(10/13。’Geograph。Bot。’page863。),ontheotherhand——andtherecannotbeamorecapablejudge——advanceswhatheconsiderssufficientevidenceoftheorange(hedoubtswhetherthebitterandsweetkindsarespecificallydistinct),thelemon,andcitron,havingbeenfoundwild,andconsequentlythattheyaredistinct。HementionstwootherformscultivatedinJapanandJava,whichheranksundoubtedspecies;hespeaksrathermoredoubtfullyabouttheshaddock,whichvariesmuch,andhasnotbeenfoundwild;andfinallyheconsiderssomeforms,suchasAdam’sappleandthebergamotte,asprobablyhybrids。

  Ihavebrieflyabstractedtheseopinionsforthesakeofshowingthosewhohaveneverattendedtosuchsubjects,howperplexingtheyare。Itwould,therefore,beuselessformypurposetogiveasketchoftheconspicuousdifferencesbetweentheseveralforms。Besidestheever—recurrentdifficultyofdeterminingwhetherformsfoundwildaretrulyaboriginalorareescapedseedlings,manyoftheforms,whichmustberankedasvarieties,transmittheircharactersalmostperfectlybyseed。Sweetandbitterorangesdifferinnoimportantrespectexceptintheflavouroftheirfruit,butGallesio(10/14。’TeoriadellaRiproduzione’pages52—57。)

  ismostemphaticthatbothkindscanbepropagatedbyseedwithabsolutecertainty。Consequently,inaccordancewithhissimplerule,heclassesthemasdistinctspecies;ashedoessweetandbitteralmonds,thepeachandnectarine,etc。Headmits,however,thatthesoft—shelledpine—treeproducesnotonlysoft—shelledbutsomehard—shelledseedlings,sothatalittlegreaterforceinthepowerofinheritancewould,accordingtothisrule,raiseasoft—shelledpine—treeintothedignityofanaboriginallycreatedspecies。ThepositiveassertionmadebyMacfayden(10/15。Hooker’Bot。Misc。’volume1page302;volume2page111。)thatthepipsofsweetorangesproducedinJamaica,accordingtothenatureofthesoilinwhichtheyaresown,eithersweetorbitteroranges,isprobablyanerror;forM。

  Alph。DeCandolleinformsmethatsincethepublicationofhisgreatworkhehasreceivedaccountsfromGuiana,theAntilles,andMauritius,thatinthesecountriessweetorangesfaithfullytransmittheircharacter。Gallesiofoundthatthewillow—leafedandtheLittleChinaorangesreproducedtheirproperleavesandfruit;buttheseedlingswerenotquiteequalinmerittotheirparents。Thered—fleshedorange,ontheotherhand,failstoreproduceitself。Gallesioalsoobservedthattheseedsofseveralothersingularvarietiesallreproducedtreeshavingapeculiarphysiognomy,partlyresemblingtheirparent—forms。Icanadduceanothercase:themyrtleleavedorangeisrankedbyallauthorsasavariety,butisverydistinctingeneralaspect:inmyfather’sgreenhouse,duringmanyyears,itrarelyyieldedanyfruit,butatlastproducedone;andatreethusraisedwasidenticalwiththeparent—form。

  Anotherandmoreseriousdifficultyindeterminingtherankoftheseveralformsisthat,accordingtoGallesio(10/16。’TeoriadellaRiproduzione’

  page53。)theylargelyintercrosswithoutartificialaid;thushepositivelystatesthatseedstakenfromlemon—trees(C。lemonum)growingmingledwiththecitron(C。medica),whichisgenerallyconsideredasadistinctspecies,producedagraduatedseriesofvarietiesbetweenthesetwoforms。Again,anAdam’sapplewasproducedfromtheseedofasweetorange,whichgrewclosetolemonsandcitrons。Butsuchfactshardlyaidusindeterminingwhethertoranktheseformsasspeciesorvarieties;foritisnowknownthatundoubtedspeciesofVerbascum,Cistus,Primula,Salix,etc。,frequentlycrossinastateofnature。Ifindeeditwereprovedthatplantsoftheorangetriberaisedfromthesecrosseswereevenpartiallysterile,itwouldbeastrongargumentinfavouroftheirrankasspecies。Gallesioassertsthatthisisthecase;buthedoesnotdistinguishbetweensterilityfromhybridismandfromtheeffectsofculture;andhealmostdestroystheforceofthisstatementbyanother(10/17。Gallesio’TeoriadellaRiproduzione’page69。)namely,thatwhenheimpregnatedtheflowersofthecommonorangewiththepollentakenfromundoubtedVARIETIESoftheorange,monstrousfruitswereproduced,whichincluded\"littlepulp,andhadnoseeds,orimperfectseeds。\"

  Inthistribeofplantswemeetwithinstancesoftwohighlyremarkablefactsinvegetablephysiology:Gallesio(10/18。Ibidpage67。)impregnatedanorangewithpollenfromalemon,andthefruitborneonthemothertreehadaraisedstripeofpeellikethatofalemonbothincolourandtaste,butthepulpwaslikethatofanorangeandincludedonlyimperfectseeds。

  Thepossibilityofpollenfromonevarietyorspeciesdirectlyaffectingthefruitproducedbyanothervarietyofspecies,isasubjectwhichI

  shallfullydiscussinthefollowingchapter。

  Thesecondremarkablefactis,thattwosupposedhybrids(10/19。Gallesio’TeoriadellaRiproduzione’pages75,76。)(fortheirhybridnaturewasnotascertained),betweenanorangeandeitheralemonorcitron,producedonthesametreeleaves,flowers,andfruitofbothpureparent—forms,aswellasofamixedorcrossednature。Abudtakenfromanyoneofthebranchesandgraftedonanothertreeproduceseitheroneofthepurekindsoracapricioustreereproducingthethreekinds。Whetherthesweetlemon,whichincludeswithinthesamefruitsegmentsofdifferentlyflavouredpulp(10/20。’Gardener’sChronicle’1841page613。),isananalogouscase,I

  knownot。ButtothissubjectIshallhavetorecur。

  IwillconcludebygivingfromA。Risso(10/21。’AnnalesduMuseum’tome20

  page188。)ashortaccountofaverysingularvarietyofthecommonorange。

  Itisthe\"citrusaurantiumfructuvariabili,\"whichontheyoungshootsproducesrounded—ovalleavesspottedwithyellow,borneonpetioleswithheart—shapedwings;whentheseleavesfalloff,theyaresucceededbylongerandnarrowerleaves,withundulatedmargins,ofapale—greencolourembroideredwithyellow,borneonfootstalkswithoutwings。Thefruitwhilstyoungispear—shaped,yellow,longitudinallystriated,andsweet;

  butasitripens,itbecomesspherical,ofareddish—yellow,andbitter。

  PEACHANDNECTARINE(Amygdaluspersica)。

  Thebestauthoritiesarenearlyunanimousthatthepeachhasneverbeenfoundwild。ItwasintroducedfromPersiaintoEuropealittlebeforetheChristianera,andatthisperiodfewvarietiesexisted。Alph。DeCandolle(10/22。’Geograph。Bot。’page882。),fromthefactofthepeachnothavingspreadfromPersiaatanearlierperiod,andfromitsnothavingpureSanscritorHebrewnames,believesthatitisnotanaboriginalofWesternAsia,butcamefromtheterraincognitaofChina。Thesupposition,however,thatthepeachisamodifiedalmondwhichacquireditspresentcharacteratacomparativelylateperiod,would,Ipresume,accountforthesefacts;onthesameprinciplethatthenectarine,theoffspringofthepeach,hasfewnativenames,andbecameknowninEuropeatastilllaterperiod。

  (FIGURE42。—PEACHANDALMONDSTONES,ofnaturalsize,viewededgeways。1。

  CommonEnglishpeach。2。Double,crimson—flowered,ChinesePeach。3。

  ChineseHoneyPeach。4。EnglishAlmond。5。BarcelonaAlmond。6。MalagaAlmond。7。Soft—shelledFrenchAlmond。8。SmyrnaAlmond。)

  AndrewKnight(10/23。’TransactionsofHort。Soc。’volume3page1andvolume4page396andnotetopage370。Acoloureddrawingisgivenofthishybrid。),fromfindingthataseedling—tree,raisedfromasweetalmondfertilisedbythepollenofapeach,yieldedfruitquitelikethatofapeach,suspectedthatthepeach—treeisamodifiedalmond;andinthishehasbeenfollowedbyvariousauthors。(10/24。’Gardener’sChronicle’1856

  page532。Awriter,itmaybepresumedDr。Lindley,remarksontheperfectserieswhichmaybeformedbetweenthealmondandthepeach。Anotherhighauthority,Mr。Rivers,whohashadsuchwideexperience,stronglysuspects(’Gardener’sChronicle’1863page27)thatpeaches,iflefttoastateofnature,wouldinthecourseoftimeretrogradeintothick—fleshedalmonds。)

  Afirst—ratepeach,almostglobularinshape,formedofsoftandsweetpulp,surroundingahard,muchfurrowed,andslightlyflattenedstone,certainlydiffersgreatlyfromanalmond,withitssoft,slightlyfurrowed,muchflattened,andelongatedstone,protectedbyatough,greenishlayerofbitterflesh。Mr。Bentham(10/25。’JournalofHort。Soc。’volume9page168。)hasparticularlycalledattentiontothestoneofthealmondbeingsomuchmoreflattenedthanthatofthepeach。Butintheseveralvarietiesofthealmond,thestonediffersgreatlyinthedegreetowhichitiscompressed,insize,shape,strength,andinthedepthofthefurrows,asmaybeseeninfigure42(Nos。4to8)ofsuchkindsasIhavebeenabletocollect。Withpeach—stonesalso(Nos。1to3)thedegreeofcompressionandelongationisseentovary;sothatthestoneoftheChineseHoney—peach(No。3)ismuchmoreelongatedandcompressedthanthatofthe(No。8)

  Smyrnaalmond。Mr。Rivers,ofSawbridgeworth,towhomIamindebtedforsomeofthespecimensabovefigured,andwhohashadsuchgreathorticulturalexperience,hascalledmyattentiontoseveralvarietieswhichconnectthealmondandthepeach。InFrancethereisavarietycalledthePeach—Almond,whichMr。Riversformerlycultivated,andwhichiscorrectlydescribedinaFrenchcatalogueasbeingovalandswollen,withtheaspectofapeach,includingahardstonesurroundedbyafleshycovering,whichissometimeseatable。(10/26。Whetherthisisthesamevarietyasonelatelymentioned(’Gardener’sChronicle’1865page1154)byM。Carriereunderthenameofpersicaintermedia,Iknownot;thisvarietyissaidtobeintermediateinnearlyallitscharactersbetweenthealmondandpeach;itproducesduringsuccessiveyearsverydifferentkindsoffruit。)AremarkablestatementbyM。Luizethasrecentlyappearedinthe’RevueHorticole’(10/27。Quotedin’Gardener’sChronicle’1866page800。),namely,thataPeach—almond,graftedonapeach,bore,during1863and1864

  almondsalone,butin1865boresixpeachesandnoalmonds。M。Carriere,incommentingonthisfact,citesthecaseofadouble—floweredalmondwhich,afterproducingduringseveralyearsalmonds,suddenlyborefortwoyearsinsuccessionsphericalfleshypeach—likefruits,butin1865revertedtoitsformerstateandproducedlargealmonds。

  Again,asIhearfromMr。Rivers,thedouble—floweringChinesepeachesresemblealmondsintheirmannerofgrowthandintheirflowers;thefruitismuchelongatedandflattened,withthefleshbothbitterandsweet,butnotuneatable,anditissaidtobeofbetterqualityinChina。Fromthisstageonesmallstepleadsustosuchinferiorpeachesasareoccasionallyraisedfromseed。Forinstance,Mr。Riverssowedanumberofpeach—stonesimportedfromtheUnitedStates,wheretheyarecollectedforraisingstocks,andsomeofthetreesraisedbyhimproducedpeacheswhichwereverylikealmondsinappearance,beingsmallandhard,withthepulpnotsofteningtillverylateintheautumn。VanMons(10/28。Quotedin’JournaldeLaSoc。Imp。d’Horticulture’1855page238。)alsostatesthatheonceraisedfromapeach—stoneapeachhavingtheaspectofawildtree,withfruitlikethatofthealmond。Frominferiorpeaches,suchasthesejustdescribed,wemaypassbysmalltransitions,throughclingstonesofpoorquality,toourbestandmostmeltingkinds。Fromthisgradation,fromthecasesofsuddenvariationaboverecorded,andfromthefactthatthepeachhasnotbeenfoundwild,itseemstomebyfarthemostprobableview,thatthepeachisthedescendantofthealmond,improvedandmodifiedinamarvellousmanner。

  Onefact,however,isopposedtothisconclusion。Ahybrid,raisedbyKnightfromthesweetalmondbythepollenofthepeach,producedflowerswithlittleornopollen,yetborefruit,havingbeenapparentlyfertilisedbyaneighbouringnectarine。Anotherhybrid,fromasweetalmondbythepollenofanectarine,producedduringthefirstthreeyearsimperfectblossoms,butafterwardsperfectflowerswithanabundanceofpollen。Ifthisslightdegreeofsterilitycannotbeaccountedforbytheyouthofthetrees(andthisoftencauseslessenedfertility),orbythemonstrousstateoftheflowers,orbytheconditionstowhichthetreeswereexposed,thesetwocaseswouldaffordagoodargumentagainstthepeachbeingthedescendantofthealmond。

  Whetherornotthepeachhasproceededfromthealmond,ithascertainlygivenrisetonectarines,orsmoothpeaches,astheyarecalledbytheFrench。Mostofthevarieties,bothofthepeachandnectarine,reproducethemselvestrulybyseed。Gallesio(10/29。’TeoriadellaRiproduzioneVegetale’1816page86。)sayshehasverifiedthiswithrespecttoeightracesofthepeach。Mr。Rivers(10/30。’Gardener’sChronicle’1862page1195。)hasgivensomestrikinginstancesfromhisownexperience,anditisnotoriousthatgoodpeachesareconstantlyraisedinNorthAmericafromseed。ManyoftheAmericansub—varietiescometrueornearlytruetotheirkind,suchasthewhite—blossom,severaloftheyellow—fruitedfreestonepeaches,thebloodclingstone,theheath,andthelemonclingstone。Ontheotherhand,aclingstonepeachhasbeenknowntogiverisetoafreestone。

  (10/31。Mr。Rivers’Gardener’sChronicle’1859page774。)InEnglandithasbeennoticedthatseedlingsinheritfromtheirparentsflowersofthesamesizeandcolour。Somecharacters,however,contrarytowhatmighthavebeenexpected,oftenarenotinherited;suchasthepresenceandformoftheglandsontheleaves。(10/32。Downing’TheFruitsofAmerica’1845pages475,489,492,494,496。SeealsoF。Michaux’TravelsinN。America’

  Englishtranslationpage228。ForsimilarcasesinFranceseeGodron’Del’Espece’tome2page97。)Withrespecttonectarines,bothclingandfreestonesareknowninNorthAmericatoreproducethemselvesbyseed。

  (10/33。Brickell’Nat。Hist。ofN。Carolina’page102andDowning’FruitTrees’page505。)InEnglandthenewwhitenectarinewasaseedlingoftheoldwhite,andMr。Rivers(10/34。’Gardener’sChronicle’1862page1196。)

  hasrecordedseveralsimilarcases。Fromthisstrongtendencytoinheritance,whichbothpeachandnectarinetreesexhibit,——fromcertainslightconstitutionaldifferences(10/35。Thepeachandnectarinedonotsucceedequallywellinthesomesoil:seeLindley’Horticulture’page351。)intheirnature,——andfromthegreatdifferenceintheirfruitbothinappearanceandflavour,itisnotsurprising,notwithstandingthatthetreesdifferinnootherrespectsandcannotevenbedistinguished,asIaminformedbyMr。Rivers,whilstyoung,thattheyhavebeenrankedbysomeauthorsasspecificallydistinct。Gallesiodoesnotdoubtthattheyaredistinct;evenAlph。DeCandolledoesnotappearperfectlyassuredoftheirspecificidentity:andaneminentbotanisthasquiterecently(10/36。

  Godron’Del’Espece’tome21859page97。)maintainedthatthenectarine\"probablyconstitutesadistinctspecies。\"

  Henceitmaybeworthwhiletogivealltheevidenceontheoriginofthenectarine。Thefactsinthemselvesarecurious,andwillhereafterhavetobereferredtowhentheimportantsubjectofbud—variationisdiscussed。Itisasserted(10/37。’Transact。Hort。Soc。’volume6page394。)thattheBostonnectarinewasproducedfromapeach—stone,andthisnectarinereproduceditselfbyseed。(10/38。Downing’s’FruitTrees’page502。)Mr。

  Riversstates(10/39。’Gardener’sChronicle’1862page1195。)thatfromstonesofthreedistinctvarietiesofthepeachheraisedthreevarietiesofnectarine;andinoneofthesecasesnonectarinegrewneartheparentpeach—tree。InanotherinstanceMr。Riversraisedanectarinefromapeach,andinthesucceedinggenerationanothernectarinefromthisnectarine。

  (10/40。’JournalofHorticulture’February5,1866page102。)Othersuchinstanceshavebeencommunicatedtome,buttheyneednotbegiven。Oftheconversecase,namely,ofnectarine—stonesyieldingpeach—trees(bothfreeandclingstones),wehavesixundoubtedinstancesrecordedbyMr。Rivers;

  andintwooftheseinstancestheparentnectarineshadbeenseedlingsfromothernectarines。(10/41。Mr。Riversin’Gardener’s’Chronicle’1859page774,1862page1195;1865page1059;and’JournalofHort。’1866page102。)

  Withrespecttothemorecuriouscaseoffull—grownpeach—treessuddenlyproducingnectarinesbybud—variation(orsportsastheyarecalledbygardeners),theevidenceissuperabundant;thereisalsogoodevidenceofthesametreeproducingbothpeachesandnectarines,orhalf—and—halffruit;bythistermImeanafruitwiththeone—halfaperfectpeach,andtheotherhalfaperfectnectarine。

  PeterCollinsonin1741recordedthefirstcaseofapeach—treeproducinganectarine(10/42。’CorrespondenceofLinnaeus’1821pages7,8,70。)andin1766headdedtwootherinstances。Inthesamework,theeditor,SirJ。E。

  Smith,describesthemoreremarkablecaseofatreeinNorfolkwhichusuallyborebothperfectnectarinesandperfectpeaches;butduringtwoseasonssomeofthefruitwerehalfandhalfinnature。

  Mr。Salisburyin1808(10/43。’Transact。Hort。Soc。’volume1page103。)

  recordssixothercasesofpeach—treesproducingnectarines。Threeofthevarietiesarenamed;viz。,theAlberge,BelleChevreuse,andRoyalGeorge。

  Thislattertreeseldomfailedtoproducebothkindsoffruit。Hegivesanothercaseofahalf—and—halffruit。

  AtRadfordinDevonshire(10/44。Loudon’s’Gardener’sMag。’1826volume1。

  page471。)aclingstonepeach,purchasedastheChancellor,wasplantedin1815,andin1824,afterhavingpreviouslyproducedpeachesalone,boreononebranchtwelvenectarines;in1825thesamebranchyieldedtwenty—sixnectarines,andin1826thirty—sixnectarines,togetherwitheighteenpeaches。Oneofthepeacheswasalmostassmoothononesideasanectarine。Thenectarineswereasdarkas,butsmallerthan,theElruge。

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