第62章
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  CHARLESDARWINTOJ。D。HOOKER。

  Down,June10[1864]……IhavenowreadtwoGermanbooks,andallIbelievethathasbeenwrittenonclimbers,andithasstirredmeuptofindthatIhaveagooddealofnewmatter。Itisstrange,butIreallythinknoonehasexplainedsimpletwiningplants。Thesebookshavestirredmeup,andmademewishforplantsspecifiedinthem。Ishallbeverygladofthoseyoumention。

  IhavewrittentoVeitchforyoungNepenthesandVanilla(whichIbelievewillturnoutagrandcase,thougharootcreeper),ifIcannotbuyyoungVanillaIwillaskyou。Ihaveorderedaleaf—climbingfern,Lygodium。

  Allthisworkaboutclimberswouldhurtmyconscience,didIthinkIcoulddoharderwork。(Hewasmuchoutofhealthatthistime。)

  [Hecontinuedhisobservationsonclimbingplantsduringtheprolongedillnessfromwhichhesufferedintheautumnof1863,andinthefollowingspring。HewrotetoSirJ。D。Hooker,apparentlyinMarch1864:——

  \"ForseveraldaysIhavebeendecidedlybetter,andwhatIlaymuchstresson(whateverdoctorssay),mybrainfeelsfarstronger,andIhavelostmanydreadfulsensations。Thehot—houseissuchanamusementtome,andmyamusementIowetoyou,asmydelightistolookatthemanyoddleavesandplantsfromKew……TheonlyapproachtoworkwhichIcandoistolookattendrilsandclimbers,thisdoesnotdistressmyweakenedbrain。AskOlivertolookovertheenclosedqueries(anddoyoulook)andamuseabroken—downbrothernaturalistbyansweringanywhichhecan。Ifyoueverloungethroughyourhouses,remembermeandclimbingplants。\"

  OnOctober29,1864,hewrotetoDr。Gray:——

  \"Ihavenotbeenabletoresistdoingalittlemoreatyourgodchild,myclimbingpaper,orratherinsizelittlebook,whichbyJoveIwillhavecopiedout,elseIshallneverstop。Thishasbeennewsortofworkforme,andIhavebeenpleasedtofindwhatacapitalguideforobservationsafullconvictionofthechangeofspeciesis。\"

  OnJanuary19,1865,hewrotetoSirJ。D。Hooker:——

  \"Itisworkinghours,butIamtryingtotakeaday’sholiday,forI

  finishedanddespatchedyesterdaymyclimbingpaper。ForthelasttendaysIhavedonenothingbutcorrectrefractorysentences,andIloathethewholesubject。\"

  AlettertoDr。Gray,April9,1865,hasawordortwoonthesubject:——

  \"Ihavebeguncorrectingproofsofmypaperon’ClimbingPlants。’I

  supposeIshallbeabletosendyouacopyinfourorfiveweeks。Ithinkitcontainsagooddealnewandsomecuriouspoints,butitissofearfullylong,thatnoonewilleverreadit。If,however,youdonotSKIMthroughit,youwillbeanunnaturalparent,foritisyourchild。\"

  Dr。Graynotonlyreaditbutapprovedofit,tomyfather’sgreatsatisfaction,asthefollowingextractsshow:——

  \"IwasmuchpleasedtogetyourletterofJuly24th。NowthatIcandonothing,Imaunderoveroldsubjects,andyourapprobationofmyclimbingpapergivesmeVERYgreatsatisfaction。ImademyobservationswhenI

  coulddonothingelseandmuchenjoyedit,butalwaysdoubtedwhethertheywereworthpublishing。IdemurtoitsnotbeingnecessarytoexplainindetailaboutthespiresinCAUGHTtendrilsrunninginoppositedirections;

  forthefactforalongtimeconfoundedme,andIhavefounditdifficultenoughtoexplainthecausetotwoorthreepersons。\"(August15,1865。)

  \"Ireceivedyesterdayyourarticle(IntheSeptembernumberof’Silliman’sJournal,’concludedintheJanuarynumber,1866。)onclimbers,andithaspleasedmeinanextraordinaryandevensillymanner。Youpaymeasuperbcompliment,andasIhavejustsaidtomywife,IthinkmyfriendsmustperceivethatIlikepraise,theygivemesuchheartydoses。Ialwaysadmireyourskillinreviewsorabstracts,andyouhavedonethisarticleexcellentlyandgiventhewholeessenceofmypaper……IhavehadaletterfromagoodZoologistinS。Brazil,F。Muller,whohasbeenstirreduptoobserveclimbersandgivesmesomecuriouscasesofBRANCH—climbers,inwhichbranchesareconvertedintotendrils,andthencontinuetogrowandthrowoutleavesandnewbranches,andthenlosetheirtendrilcharacter。\"

  (October1865。)

  ThepaperonClimbingPlantswasrepublishedin1875,asaseparatebook。

  Theauthorhadbeenunabletogivehiscustomaryamountofcaretothestyleoftheoriginalessay,owingtothefactthatitwaswrittenduringaperiodofcontinuedill—health,anditwasnowfoundtorequireagreatdealofalteration。HewrotetoSirJ。D。Hooker(March3,1875):\"Itisluckyforauthorsingeneralthattheydonotrequiresuchdreadfulworkinmerelylickingwhattheywriteintoshape。\"AndtoMr。MurrayinSeptemberhewrote:\"Thecorrectionsareheavyin’ClimbingPlants,’andyetI

  deliberatelywentovertheMS。andoldsheetsthreetimes。\"ThebookwaspublishedinSeptember1875,aneditionof1500copieswasstruckoff;theeditionsoldfairlywell,and500additionalcopieswereprintedinJuneofthefollowingyear。]

  INSECTIVOROUSPLANTS。

  [Inthesummerof1860hewasstayingatthehouseofhissister—in—law,MissWedgwood,inAshdownForest,whencehewrote(July29,1860),toSirJosephHooker;——

  \"LatterlyIhavedonenothinghere;butatfirstIamusedmyselfwithafewobservationsontheinsect—catchingpowerofDrosera;andImustconsultyousometimewhethermy’twaddle’isworthcommunicatingtotheLinneanSociety。\"

  InAugusthewrotetothesamefriend:——

  \"IwillgratefullysendmynotesonDroserawhencopiedbymycopier:thesubjectamusedmewhenIhadnothingtodo。\"

  Hehasdescribedinthe’Autobiography’(volumei。),thegeneralnatureoftheseearlyexperiments。Henoticedinsectsstickingtotheleaves,andfindingthatflies,etc。,placedontheadhesiveglandswereheldfastandembraced,hesuspectedthattheleaveswereadaptedtosupplynitrogenousfoodtotheplant。Hethereforetriedtheeffectontheleavesofvariousnitrogenousfluids——withresultswhich,asfarastheywent,verifiedhissurmise。InSeptember,1860,hewrotetoDr。Gray:——

  \"IhavebeeninfinitelyamusedbyworkingatDrosera:themovementsarereallycurious;andthemannerinwhichtheleavesdetectcertainnitrogenouscompoundsismarvellous。Youwilllaugh;butitis,atpresent,myfullbelief(afterendlessexperiments)thattheydetect(andmoveinconsequenceof)the1/2880partofasinglegrainofnitrateofammonia;butthemuriateandsulphateofammoniabothertheirchemicalskill,andtheycannotmakeanythingofthenitrogeninthesesalts!I

  beganthisworkonDroserainrelationtoGRADATIONasthrowinglightonDionaea。\"

  LaterintheautumnhewasagainobligedtoleavehomeforEastbourne,wherehecontinuedhisworkonDrosera。Theworkwassonewtohimthathefoundhimselfindifficultiesinthepreparationofsolutions,andbecamepuzzledoverfluidandsolidounces,etc。etc。Toafriend,thelateMr。

  E。Cresy,whocametohishelpinthematterofweightsandmeasures,hewrotegivinganaccountoftheexperiments。Theextract(November2,1860)

  whichfollowsillustratesthealmostsuperstitiousprecautionsheoftenappliedtohisresearches:——

  \"GenerallyIhavescrutinisedeveryglandandhairontheleafbeforeexperimenting;butitoccurredtomethatImightinsomewayaffecttheleaf;thoughthisisalmostimpossible,asIscrutinisedwithequalcarethosethatIputintodistilledwater(thesamewaterbeingusedfordissolvingthecarbonateofammonia)。Ithencutofffourleaves(nottouchingthemwithmyfingers),andputtheminplainwater,andfourotherleavesintotheweaksolution,andafterleavingthemforanhourandahalf,Iexaminedeveryhaironalleightleaves;nochangeonthefourinwater;everyglandandhairaffectedinthoseinammonia。

  \"Ihadmeasuredthequantityofweaksolution,andIcountedtheglandswhichhadabsorbedtheammonia,andwereplainlyaffected;theresultconvincedmethateachglandcouldnothaveabsorbedmorethan1/64000or1/65000ofagrain。Ihavetriednumbersofotherexperimentsallpointingtothesameresult。Someexperimentsleadmetobelievethatverysensitiveleavesareactedonbymuchsmallerdoses。Reflecthowlittleammoniaaplantcangetgrowingonpoorsoil——yetitisnourished。Thereallysurprisingpartseemstomethattheeffectshouldbevisible,andnotunderveryhighpower;foraftertryingahighpower,Ithoughtitwouldbesafernottoconsideranyeffectwhichwasnotplainlyvisibleunderatwo—thirdsobjectglassandmiddleeye—piece。Theeffectwhichthecarbonateofammoniaproducesisthesegregationofthehomogeneousfluidinthecellsintoacloudofgranulesandcolourlessfluid;andsubsequentlythegranulescoalesceintolargermasses,andforhourshavetheoddestmovements——coalescing,dividing,coalescingadinfinitum。Idonotknowwhetheryouwillcarefortheseill—writtendetails;but,asyouasked,IamsureIamboundtocomply,afteralltheverykindandgreattroublewhichyouhavetaken。\"

  OnhisreturnhomehewrotetoSirJ。D。Hooker(November21,1860):——

  \"IhavebeenworkinglikeamadmanatDrosera。Hereisafactforyouwhichiscertainasyoustandwhereyouare,thoughyouwon’tbelieveit,thatabitofhair1/78000ofonegraininweightplacedongland,willcauseONEofthegland—bearinghairsofDroseratocurveinwards,andwillaltertheconditionofthecontentsofeverycellinthefoot—stalkofthegland。\"

  AndafewdayslatertoLyell:——

  \"IwillandmustfinishmyDroseraMS。,whichwilltakemeaweek,for,atthepresentmoment,IcaremoreaboutDroserathantheoriginofallthespeciesintheworld。ButIwillnotpublishonDroseratillnextyear,forIamfrightenedandastoundedatmyresults。Ideclareitisacertainfact,thatoneorganissosensitivetotouch,thataweightseventy—eighttimeslessthanthat,viz。,1/1000ofagrain,whichwillmovethebestchemicalbalance,sufficestocauseaconspicuousmovement。Isitnotcuriousthataplantshouldbefarmoresensitivetothetouchthananynerveinthehumanbody?YetIamperfectlysurethatthisistrue。WhenIamonmyhobby—horse,Inevercanresisttellingmyfriendshowwellmyhobbygoes,soyoumustforgivetherider。\"

  Theworkwascontinued,asaholidaytask,atBournemouth,wherehestayedduringtheautumnof1862。Thediscussioninthefollowingletteron\"nervousmatter\"inDroseraisofinterestinrelationtorecentresearchesonthecontinuityofprotoplasmfromcelltocell:]

  CHARLESDARWINTOJ。D。HOOKER。

  CliffCottage,Bournemouth。

  September26[1862]。

  MydearHooker,Donotreadthistillyouhaveleisure。Ifthatblessedmomentevercomes,Ishouldbeverygladtohaveyouropiniononthesubjectofthisletter。

  IamledtotheopinionthatDroseramusthavediffusedmatterinorganicconnection,closelyanalogoustothenervousmatterofanimals。Whentheglandsofoneofthepapillaeortentacles,initsnaturalpositionissuppliedwithnitrogenisedfluidandcertainotherstimulants,orwhenloadedwithanextremelyslightweight,orwhenstruckseveraltimeswithaneedle,thepedicelbendsnearitsbaseinunderoneminute。Thesevariedstimulantsareconveyeddownthepedicelbysomemeans;itcannotbevibration,fordropsoffluidputonquitequietlycausethemovement;itcannotbeabsorptionofthefluidfromcelltocell,forIcanseetherateofabsorption,whichthoughquick,isfarslower,andinDionaeathetransmissionisinstantaneous;analogyfromanimalswouldpointtotransmissionthroughnervousmatter。Reflectingontherapidpowerofabsorptionintheglands,theextremesensibilityofthewholeorgan,andtheconspicuousmovementcausedbyvariedstimulants,Ihavetriedanumberofsubstanceswhicharenotcausticorcorrosive,……butmostofwhichareknowntohavearemarkableactiononthenervousmatterofanimals。Youwillseetheresultsintheenclosedpaper。Asthenervousmatterofdifferentanimalsaredifferentlyactedonbythesamepoisons,onewouldnotexpectthesameactiononplantsandanimals;onlyifplantshavediffusednervousmatter,somedegreeofanalogousaction。Andthisispartiallythecase。Consideringtheseexperiments,togetherwiththepreviouslymaderemarksonthefunctionsoftheparts,Icannotavoidtheconclusion,thatDroserapossessesmatteratleastinsomedegreeanalogousinconstitutionandfunctiontonervousmatter。Nowdotellmewhatyouthink,asfarasyoucanjudgefrommyabstract;ofcoursemanymoreexperimentswouldhavetobetried;butinformeryearsItriedonthewholeleaf,insteadofonseparateglands,anumberofinnocuous(Thislineofinvestigationmadehimwishforinformationontheactionofpoisonsonplants;asinmanyothercasesheappliedtoProfessorOliver,andinreferencetotheresultwrotetoHooker:\"PraythankOliverheartilyforhisheapofreferencesonpoisons。\")substances,suchassugar,gum,starch,etc。,andtheyproducednoeffect。Youropinionwillaidmeindecidingsomefutureyearingoingonwiththissubject。Ishouldnothavethoughtitworthattempting,butIhadnothingonearthtodo。

  MydearHooker,Yoursverysincerely,CH。DARWIN。

  P。S。——WereturnhomeonMonday28th。ThankHeaven!

  [Alongbreaknowensuedinhisworkoninsectivorousplants,anditwasnottill1872thatthesubjectseriouslyoccupiedhimagain。ApassageinalettertoDr。AsaGray,writtenin1863or1864,shows,however,thatthequestionwasnotaltogetherabsentfromhismindintheinterim:——

  \"DependonityouareunjustonthemeritsofmybelovedDrosera;itisawonderfulplant,orratheramostsagaciousanimal。IwillstickupforDroseratothedayofmydeath。HeavenknowswhetherIshalleverpublishmypileofexperimentsonit。\"

  Henotesinhisdiarythatthelastproofofthe’ExpressionoftheEmotions’wasfinishedonAugust22,1872,andthathebegantoworkonDroseraonthefollowingday。]

  CHARLESDARWINTOASAGRAY。

  [Sevenoaks],October22[1872]……IhaveworkedprettyhardforfourorfiveweeksonDrosera,andthenbrokedown;sothatwetookahousenearSevenoaksforthreeweeks(whereI

  nowam)togetcompleterest。Ihaveverylittlepowerofworkingnow,andmustputofftherestoftheworkonDroseratillnextspring,asmyplantsaredying。Itisanendlesssubject,andImustcutitshort,andforthisreasonshallnotdomuchonDionaea。ThepointwhichhasinterestedmemostistracingtheNERVES!whichfollowthevascularbundles。Byaprickwithasharplancetatacertainpoint,Icanparalyseone—halftheleaf,sothatastimulustotheotherhalfcausesnomovement。Itisjustlikedividingthespinalmarrowofafrog:——nostimuluscanbesentfromthebrainoranteriorpartofthespinetothehindlegs;butiftheselatterarestimulated,theymovebyreflexaction。Ifindmyoldresultsabouttheastonishingsensitivenessofthenervoussystem(!?)ofDroseratovariousstimulantsfullyconfirmedandextended……

  [HisworkondigestioninDroseraandotherpointsinthephysiologyoftheplantsoonledhimintoregionswherehisknowledgewasdefective,andheretheadviceandassistancewhichhereceivedfromDr。BurdonSandersonwasofmuchvalue:]

  CHARLESDARWINTOJ。BURDONSANDERSON。

  Down,July25,1873。

  MydearDr。Sanderson,IshouldliketotellyoualittleaboutmyrecentworkwithDrosera,toshowthatIhaveprofitedbyyoursuggestions,andtoaskaquestionortwo。

  1。ItisreallybeautifulhowquicklyandwellDroseraandDionaeadissolvelittlecubesofalbumenandgelatine。Ikeptthesamesizedcubesonwetmossforcomparison。WhenyouwerehereIforgotthatIhadtriedgelatine,butalbumenisfarbetterforwatchingitsdissolutionandabsorption。Franklandhastoldmehowtotestinaroughwayforpepsin;

  andintheautumnhewilldiscoverwhatacidthedigestivejuicecontains。

  2。Adecoctionofcabbage—leavesandgreenpeascausesasmuchinflectionasaninfusionofrawmeat;adecoctionofgrassislesspowerful。ThoughIhearthatthechemiststrytoprecipitateallalbumenfromtheextractofbelladonna,Ithinktheymustfail,astheextractcausesinflection,whereasanewlotofatropine,aswellasthevalerianate[ofatropine],producenoeffect。

  3。Ihavebeentryingagoodmanyexperimentswithheatedwater……Shouldyounotcallthefollowingcaseoneofheatrigor?Twoleaveswereheatedto130deg,andhadeverytentaclecloselyinflected;onewastakenoutandplacedincoldwater,anditre—expanded;theotherwasheatedto145deg,andhadnottheleastpowerofre—expansion。Isnotthislattercaseheatrigor?Ifyoucaninformme,Ishouldverymuchliketohearatwhattemperaturecold—bloodedandinvertebrateanimalsarekilled。

  4。Imusttellyoumyfinalresult,ofwhichIamsure,[asto]thesensitivenessofDrosera。Imadeasolutionofonepartofphosphateofammoniabyweightto218,750ofwater;ofthissolutionIgavesomuchthataleafgot1/8000ofagrainofthephosphate。Ithencountedtheglands,andeachcouldhavegotonly1/1552000ofagrain;thisbeingabsorbedbytheglands,sufficedtocausethetentaclesbearingtheseglandstobendthroughanangleof180deg。Suchsensitivenessrequireshotweather,andcarefullyselectedyoungyetmatureleaves。Itstrikesmeasawonderfulfact。ImustaddthatItookeveryprecaution,bytryingnumerousleavesatthesametimeinthesolutionandinthesamewaterwhichwasusedformakingthesolution。

  5。Ifyoucanpersuadeyourfriendtotrytheeffectsofcarbonateofammoniaontheaggregationofthewhitebloodcorpuscles,Ishouldverymuchliketoheartheresult。

  Ihopethisletterwillnothaveweariedyou。

  Believeme,yoursverysincerely,CHARLESDARWIN。

  CHARLESDARWINTOW。THISELTONDYER。

  Down,24[December1873?]。

  MydearMr。Dyer,Ifearthatyouwillthinkmeagreatbore,butIcannotresisttellingyouthatIhavejustfoundoutthattheleavesofPinguiculapossessabeautifullyadaptedpowerofmovement。LastnightIputonarowoflittlefliesnearoneedgeoftwoYOUNGISHleaves;andafter14hourstheseedgesarebeautifullyfoldedoversoastoclasptheflies,thusbringingtheglandsintocontactwiththeuppersurfacesoftheflies,andtheyarenowsecretingcopiouslyaboveandbelowthefliesandnodoubtabsorbing。Theacidsecretionhasrundownthechannellededgeandhascollectedinthespoon—shapedextremity,wherenodoubttheglandsareabsorbingthedelicioussoup。Theleafononesidelooksjustlikethehelixofahumanear,ifyouweretostuffflieswithinthefold。Yoursmostsincerely,CH。DARWIN。

  CHARLESDARWINTOASAGRAY。

  Down,June3[1874]……IamnowhardatworkgettingmybookonDrosera&Co。readyfortheprinters,butitwilltakesometime,forIamalwaysfindingoutnewpointstoobserve。IthinkyouwillbeinterestedbymyobservationsonthedigestiveprocessinDrosera;thesecretioncontainsanacidoftheaceticseries,andsomefermentcloselyanalogousto,butnotidenticalwith,pepsin;forIhavebeenmakingalongseriesofcomparativetrials。

  NohumanbeingwillbelievewhatIshallpublishaboutthesmallnessofthedosesofphosphateofammoniawhichact……IbeganreadingtheMadagascarsquib(Adescriptionofacarnivorousplantsupposedtosubsistonhumanbeings。)quitegravely,andwhenIfounditstatedthatFelisandBosinhabitedMadagascar,Ithoughtitwasafalsestory,anddidnotperceiveitwasahoaxtillIcametothewoman……

  CHARLESDARWINTOF。C。DONDERS。(ProfessorDonders,thewell—knownphysiologistofUtrecht。)

  Down,July7,1874。

  MydearProfessorDonders,MysonGeorgewritestomethathehasseenyou,andthatyouhavebeenverykindtohim,forwhichIreturntoyoumycordialthanks。Hetellsmeonyourauthority,ofafactwhichinterestsmeinthehighestdegree,andwhichImuchwishtobeallowedtoquote。Itrelatestotheactionofonemillionthofagrainofatropineontheeye。Nowwillyoubesokind,wheneveryoucanfindalittleleisure,totellmewhetheryouyourselfhaveobservedthisfact,orbelieveitongoodauthority。Ialsowishtoknowwhatproportionbyweighttheatropineboretothewatersolution,andhowmuchofthesolutionwasappliedtotheeye。ThereasonwhyIamsoanxiousonthisheadisthatitgivessomesupporttocertainfactsrepeatedlyobservedbymewithrespecttotheactionofphosphateofammoniaonDrosera。The1/4000000ofagrainabsorbedbyaglandclearlymakesthetentaclewhichbearsthisglandbecomeinflected;andIamfullyconvincedthat1/20000000ofagrainofthecrystallisedsalt(i。e。

  containingaboutone—thirdofitsweightofwaterofcrystallisation)doesthesame。NowIamquiteunhappyatthethoughtofhavingtopublishsuchastatement。Itwillbeofgreatvaluetometobeabletogiveanyanalogousfactsinsupport。ThecaseofDroseraisallthemoreinterestingastheabsorptionofthesaltoranyotherstimulantappliedtotheglandcausesittotransmitamotorinfluencetothebaseofthetentaclewhichbearsthegland。

  Prayforgivemefortroublingyou,anddonottroubleyourselftoanswerthisuntilyourhealthisfullyre—established。

  Praybelieveme,Yoursverysincerely,CHARLESDARWIN。

  [Duringthesummerof1874hewasatworkonthegenusUtricularia,andhewrote(July16th)toSirJ。D。Hookergivingsomeaccountoftheprogressofhiswork:——

  \"IamrathergladyouhavenotbeenabletosendUtricularia,forthecommonspecieshasdrivenF。andmealmostmad。ThestructureisMOST

  complex。ThebladderscatchamultitudeofEntomostraca,andlarvaeofinsects。Themechanismforcaptureisexcellent。Butthereismuchthatwecannotunderstand。FromwhatIhaveseento—day,Istronglysuspectthatitisnecrophagous,i。e。thatitcannotdigest,butabsorbsdecayingmatter。\"

  HewasindebtedtoLadyDorothyNevillforspecimensofthecuriousUtriculariamontana,whichisnotaquaticliketheEuropeanspecies,butgrowsamongthemossanddebrisonthebranchesoftrees。Tothisspeciesthefollowingletterrefers:]

  CHARLESDARWINTOLADYDOROTHYNEVILL。

  DownSeptember18[1874]。

  DearLadyDorothyNevill,Iamsomuchobligedtoyou。IwassoconvincedthatthebladderswerewiththeleavesthatIneverthoughtofremovingthemoss,andthiswasverystupidofme。Thegreatsolidbladder—likeswellingsalmostonthesurfacearewonderfulobjects,butarenotthetruebladders。TheseI

  foundontherootsnearthesurface,anddowntoadepthoftwoinchesinthesand。Theyareastransparentasglass,from1/20to1/100ofaninchinsize,andhollow。TheyhavealltheimportantpointsofstructureofthebladdersofthefloatingEnglishspecies,andIfeltconfidentIshouldfindcapturedprey。AndsoIhavetomydelightintwobladders,withclearproofthattheyhadabsorbedfoodfromthedecayingmass。ForUtriculariaisacarrion—feeder,andnotstrictlycarnivorouslikeDrosera。

  Thegreatsolidbladder—likebodies,Ibelieve,arereservoirsofwaterlikeacamel’sstomach。AssoonasIhavemadeafewmoreobservations,I

  meantobesocruelastogiveyourplantnowater,andobservewhetherthegreatbladdersshrinkandcontainairinsteadofwater;Ishallthenalsowashallearthfromallroots,andseewhethertherearetruebladdersforcapturingsubterraneaninsectsdowntotheverybottomofthepot。Nowshallyouthinkmeverygreedy,ifIsaythatsupposingthespeciesisnotveryprecious,andyouhaveseveral,willyougivemeonemoreplant,andifso,pleasetosenditto\"OrpingtonStation,S。E。R。,tobeforwardedbyfootmessenger。\"

  IhavehardlyeverenjoyedadaymoreinmylifethanIhavethisday’swork;andthisIowetoyourLadyship’sgreatkindness。

  Theseedsareverycuriousmonsters;IfancyofsomeplantalliedtoMedicago,butIwillshowthemtoDr。Hooker。

  Yourladyship’sverygratefully,CH。DARWIN。

  CHARLESDARWINTOJ。D。HOOKER。

  Down,September30,1874。

  MydearH。,YourmagnificentpresentofAldrovandahasarrivedquitesafe。Ihaveenjoyedgreatlyagoodlookattheshutleaves,oneofwhichIcutopen。

  ItisanaquaticDionaea,whichhasacquiredsomestructuresidenticalwiththoseofUtricularia!

  IftheleavesopenandIcantransferthemopenunderthemicroscope,I

  willtrysomeexperiments,formortalmancannotresistthetemptation。IfIcannottransfer,Iwilldonothing,forotherwiseitwouldrequirehundredsofleaves。

  Youareagoodmantogivemesuchpleasure。

  Yoursaffectionately,C。DARWIN。

  [Themanuscriptof’InsectivorousPlants’wasfinishedinMarch1875。Heseemstohavebeenmorethanusuallyoppressedbythewritingofthisbook,thushewrotetoSirJ。D。HookerinFebruary:——

  \"Youaskaboutmybook,andallthatIcansayisthatIamreadytocommitsuicide;Ithoughtitwasdecentlywritten,butfindsomuchwantsrewriting,thatitwillnotbereadytogotoprintersfortwomonths,andwillthenmakeaconfoundedlybigbook。Murraywillsaythatitisnousepublishinginthemiddleofsummer,soIdonotknowwhatwillbetheupshot;butIbegintothinkthateveryonewhopublishesabookisafool。\"

  ThebookwaspublishedonJuly2nd,1875,and2700copiesweresoldoutoftheeditionof3000。]

  CHAPTER2。XIV。

  THE’POWEROFMOVEMENTINPLANTS。’

  1880。

  [Thefewsentencesintheautobiographicalchaptergivewithsufficientclearnesstheconnectionbetweenthe’PowerofMovement,’andoneoftheauthor’searlierbooks,thaton’ClimbingPlants。’Thecentralideaofthebookisthatthemovementsofplantsinrelationtolight,gravitation,etc。,aremodificationsofaspontaneoustendencytorevolveorcircumnutate,whichiswidelyinherentinthegrowingpartsofplants。

  Thisconceptionhasnotbeengenerallyadopted,andhasnottakenaplaceamongthecanonsoforthodoxphysiology。ThebookhasbeentreatedbyProfessorSachswithafewwordsofprofessorialcontempt;andbyProfessorWiesnerithasbeenhonouredbycarefulandgenerouslyexpressedcriticism。

  Mr。ThiseltonDyer(’CharlesDarwin’(’Nature’Series),page41。)haswellsaid:\"Whetherthismasterlyconceptionoftheunityofwhathashithertoseemedachaosofunrelatedphenomenawillbesustained,timealonewillshow。ButnoonecandoubttheimportanceofwhatMr。Darwinhasdone,inshowingthatforthefuturethephenomenaofplantmovementcanandindeedmustbestudiedfromasinglepointofview。\"

  Theworkwasbeguninthesummerof1877,afterthepublicationof’DifferentFormsofFlowers,’andbytheautumnhisenthusiasmforthesubjectwasthoroughlyestablished,andhewrotetoMr。Dyer:\"Iamallonfireatthework。\"Atthistimehewasstudyingthemovementsofcotyledons,inwhichthesleepofplantsistobeobservedinitssimplestform;inthefollowingspringhewastryingtodiscoverwhatusefulpurposethesesleep—movementscouldserve,andwrotetoSirJosephHooker(March25th,1878):——

  \"IthinkwehavePROVEDthatthesleepofplantsistolessentheinjurytotheleavesfromradiation。Thishasinterestedmemuch,andhascostusgreatlabour,asithasbeenaproblemsincethetimeofLinnaeus。Butwehavekilledorbadlyinjuredamultitudeofplants:N。B。——Oxaliscarnosawasmostvaluable,butlastnightwaskilled。\"

  Hislettersofthisperioddonotgiveanyconnectedaccountoftheprogressofthework。Thetwofollowingaregivenasbeingcharacteristicoftheauthor:]

  CHARLESDARWINTOW。THISELTONDYER。

  Down,June2,1878。

  MydearDyer,IremembersayingthatIshoulddieadisgracedmanifIdidnotobserveaseedlingCactusandCycas,andyouhavesavedmefromthishorriblefate,astheymovesplendidlyandnormally。ButIhavetwoquestionstoask:

  theCycasobservedwasahugeseedinabroadandveryshallowpotwithcocoa—nutfibreasIsuppose。ItwasnamedonlyCycas。WasitCycaspectinata?IsupposethatIcannotbewronginbelievingthatwhatfirstappearsabovegroundisatrueleaf,forIcanseenostemoraxis。

  Lastly,youmayrememberthatIsaidthatwecouldnotraiseOpuntianigricans;nowImustconfesstoapieceofstupidity;onedidcomeup,butmygardenerandselfstaredatit,andconcludedthatitcouldnotbeaseedlingOpuntia,butnowthatIhaveseenoneofO。basilaris,Iamsureitwas;Iobserveditonlycasually,andsawmovements,whichmakesmewishtoobservecarefullyanother。Ifyouhaveanyfruit,willMr。Lynch(Mr。

  R。I。Lynch,nowCuratoroftheBotanicGardenatCambridgewasatthistimeintheRoyalGardens,Kew。)besokindastosendonemore?

  Iamworkingawaylikeaslaveatradicles[roots]andatmovementsoftrueleaves,forIhaveprettywelldonewithcotyledons……

  ThatwasanEXCELLENTletterabouttheGardens(ThisreferstoanattempttoinducetheGovernmenttoopentheRoyalGardensatKewinthemorning。):

  Ihadhopedthattheagitationwasover。Politiciansareapoortrucklinglot,for[they]mustseethewretchedeffectsofkeepingthegardensopenalldaylong。

  Yourevertroublesomefriend,CH。DARWIN。

  CHARLESDARWINTOW。THISELTONDYER。

  4BryanstonSt。,PortmanSquare,November21[1878]。

  MydearDyer,ImustthankyouforallthewonderfultroublewhichyouhavetakenabouttheseedsofImpatiens,andonscoresofotheroccasions。Itintruthmakesmefeelashamedofmyself,andIcannothelpthinking:\"OhLord,whenheseesourbookhewillcryout,isthisallforwhichIhavehelpedsomuch!\"Inseriousness,Ihopethatwehavemadeoutsomepoints,butI

  fearthatwehavedoneverylittleforthelabourwhichwehaveexpendedonourwork。Wearehereforaweekforalittlerest,whichIneeded。

  IfIrememberright,November30th,istheanniversaryattheRoyal,andI

  fearSirJosephmustbealmostatthelastgasp。IshallbegladwhenheisnolongerPresident。

  Yoursverysincerely,CH。DARWIN。

  [Inthespringofthefollowingyear,1879。Whenhewasengagedinputtinghisresultstogether,hewrotesomewhatdespondinglytoMr。Dyer:\"Iamoverwhelmedwithmynotes,andalmosttoooldtoundertakethejobwhichI

  haveinhand——i。e。movementsofallkinds。Yetitisworsetobeidle。\"

  Lateronintheyear,whentheworkwasapproachingcompletion,hewrotetoProf。Carus(July17,1879),withrespecttoatranslation:——

  \"TogetherwithmysonFrancis,IampreparingaratherlargevolumeonthegeneralmovementsofPlants,andIthinkthatwehavemadeoutagoodmanynewpointsandviews。

  \"IfearthatourviewswillmeetagooddealofoppositioninGermany;butwehavebeenworkingveryhardforsomeyearsatthesubject。

  \"IshallbeMUCHpleasedifyouthinkthebookworthtranslating,andproof—sheetsshallbesentyou,whenevertheyareready。\"

  Intheautumnhewashardatworkonthemanuscript,andwrotetoDr。Gray(October24,1879):——

  \"Ihavewrittenaratherbigbook——moreisthepity——onthemovementsofplants,andIamnowjustbeginningtogoovertheMS。forthesecondtime,whichisahorridbore。\"

  Onlytheconcludingpartofthenextletterreferstothe’PowerofMovements’:]

  CHARLESDARWINTOA。DECANDOLLE。

  May28,1880。

  MydearSir,Iamparticularlyobligedtoyouforhavingsokindlysendmeyour’Phytographie’(Abookonthemethodsofbotanicalresearch,moreespeciallyofsystematicwork。);forifIhadmerelyseenitadvertised,I

  shouldnothavesupposedthatitcouldhaveconcernedme。Asitis,Ihavereadwithverygreatinterestaboutaquarter,butwillnotdelaylongerthankingyou。Allthatyousayseemstomeveryclearandconvincing,andasinallyourwritingsIfindalargenumberofphilosophicalremarksnewtome,andnodoubtshallfindmanymore。TheyhaverecalledmanyapuzzlethroughwhichIpassedwhenmonographingtheCirripedia;andyourbookinthosedayswouldhavebeenquiteinvaluabletome。IthaspleasedmetofindthatIhavealwaysfollowedyourplanofmakingnotesonseparatepiecesofpaper;Ikeepseveralscoresoflargeportfolios,arrangedonverythinshelvesabouttwoinchesapart,fastenedtothewallsofmystudy,andeachshelfhasitspropernameortitle;andIcanthusputatonceeverymemorandumintoitsproperplace。Yourbookwill,Iamsure,beveryusefultomanyyoungstudents,andIshallbegmysonFrancis(whointendstodevotehimselftothephysiologyofplants)toreaditcarefully。

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