Darwinian;butwhatevidenceotherthaninferentialcanfromthenatureofthecasebeadducedinsupportofthis,asIbelieve,perfectlycorrectjudgment?NoneknowbetterthantheywhoclamourfordirectevidencethattheirmasterwasrightintakingthepositionassignedtohimbyProfessorVines,thattheycannotreasonablylookforit。Withus,aswiththemselves,modificationproceedsverygradually,anditviolatesourprinciplesasmuchastheirowntoexpectvisiblepermanentprogress,inanysinglegeneration,orindeedinanynumberofgenerationsofwildspecieswhichwehaveyethadtimetoobserve。Occasionallywecanfindsuchcases,asinthatofBranchipusstagnalis,quotedbyMr。
Wallace,orinthatoftheNewZealandKeawhoseskin,IwasassuredbythelateSirJuliusvonHaast,hasalreadybeenmodifiedasaconsequenceofitschangeoffood。Herewecanshowthatinevenafewgenerationsstructureismodifiedunderchangedconditionsofexistence,butaswebelievethesecasestooccurcomparativelyrarely,soitisstillmorerarelythattheyoccurwhenandwherewecanwatchthem。Natureiseminentlyconservative,andfixityoftype,evenunderconsiderablechangeofconditions,issurelymoreimportantforthewell-beingofanyspeciesthananover-readypowerofadaptationto,itmaybe,passingchanges。Therecouldbenosteadyprogressifeachgenerationwerenotmainlyboundbythetraditionsofthosethathavegonebeforeit。Itisevolutionandnotincessantrevolutionthatbothpartiesareupholding;andthisbeingso,rapidvisiblemodificationmustbetheexception,nottherule。Ihavequoteddirectevidenceadducedbycompetentobservers,whichis,Ibelieve,sufficienttoestablishthefactthatoffspringcanbeandissometimesmodifiedbytheacquiredhabitsofaprogenitor。Iwillnowproceedtothestillmore,asitappearstome,cogentproofaffordedbygeneralconsiderations。
What,letmeask,aretheprincipalphenomenaofheredity?Theremustbephysicalcontinuitybetweenparent,orparents,andoffspring,sothattheoffspringis,asErasmusDarwinwellsaid,akindofelongationofthelifeoftheparent。
ErasmusDarwinputthemattersowellthatImayaswellgivehiswordsinfull;hewrote:-
\"Owingtotheimperfectionoflanguagetheoffspringistermedanewanimal,butisintruthabranchorelongationoftheparent,sinceapartoftheembryonanimalis,orwas,apartoftheparent,andtherefore,instrictlanguage,cannotbesaidtobeentirelynewatthetimeofitsproduction;andthereforeitmayretainsomeofthehabitsoftheparentsystem。