第8章
加入书架 A- A+
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  Darwinian;butwhatevidenceotherthaninferentialcanfromthenatureofthecasebeadducedinsupportofthis,asIbelieve,perfectlycorrectjudgment?NoneknowbetterthantheywhoclamourfordirectevidencethattheirmasterwasrightintakingthepositionassignedtohimbyProfessorVines,thattheycannotreasonablylookforit。Withus,aswiththemselves,modificationproceedsverygradually,anditviolatesourprinciplesasmuchastheirowntoexpectvisiblepermanentprogress,inanysinglegeneration,orindeedinanynumberofgenerationsofwildspecieswhichwehaveyethadtimetoobserve。Occasionallywecanfindsuchcases,asinthatofBranchipusstagnalis,quotedbyMr。

  Wallace,orinthatoftheNewZealandKeawhoseskin,IwasassuredbythelateSirJuliusvonHaast,hasalreadybeenmodifiedasaconsequenceofitschangeoffood。Herewecanshowthatinevenafewgenerationsstructureismodifiedunderchangedconditionsofexistence,butaswebelievethesecasestooccurcomparativelyrarely,soitisstillmorerarelythattheyoccurwhenandwherewecanwatchthem。Natureiseminentlyconservative,andfixityoftype,evenunderconsiderablechangeofconditions,issurelymoreimportantforthewell-beingofanyspeciesthananover-readypowerofadaptationto,itmaybe,passingchanges。Therecouldbenosteadyprogressifeachgenerationwerenotmainlyboundbythetraditionsofthosethathavegonebeforeit。Itisevolutionandnotincessantrevolutionthatbothpartiesareupholding;andthisbeingso,rapidvisiblemodificationmustbetheexception,nottherule。Ihavequoteddirectevidenceadducedbycompetentobservers,whichis,Ibelieve,sufficienttoestablishthefactthatoffspringcanbeandissometimesmodifiedbytheacquiredhabitsofaprogenitor。Iwillnowproceedtothestillmore,asitappearstome,cogentproofaffordedbygeneralconsiderations。

  What,letmeask,aretheprincipalphenomenaofheredity?Theremustbephysicalcontinuitybetweenparent,orparents,andoffspring,sothattheoffspringis,asErasmusDarwinwellsaid,akindofelongationofthelifeoftheparent。

  ErasmusDarwinputthemattersowellthatImayaswellgivehiswordsinfull;hewrote:-

  \"Owingtotheimperfectionoflanguagetheoffspringistermedanewanimal,butisintruthabranchorelongationoftheparent,sinceapartoftheembryonanimalis,orwas,apartoftheparent,andtherefore,instrictlanguage,cannotbesaidtobeentirelynewatthetimeofitsproduction;andthereforeitmayretainsomeofthehabitsoftheparentsystem。

  \"Attheearliestperiodofitsexistencetheembryonwouldseemtoconsistofalivingfilamentwithcertaincapabilitiesofirritation,sensation,volition,andassociation,andalsowithsomeacquiredhabitsorpropensitiespeculiartotheparent;theformeroftheseareincommonwithotheranimals;thelatterseemtodistinguishorproducethekindofanimal,whethermanorquadruped,withthesimilarityoffeatureorformtotheparent。\"{39}

  Thosewhoacceptevolutioninsistonunbrokenphysicalcontinuitybetweentheearliestknownlifeandourselves,sothatwebothareandarenotpersonallyidenticalwiththeunicellularorganismfromwhichwehavedescendedinthecourseofmanymillionsofyears,exactlyinthesamewayasanoctogenarianbothisandisnotpersonallyidenticalwiththemicroscopicimpregnateovumfromwhichhegrewup。Everythingbothisandisnot。Thereisnosuchthingasstrictidentitybetweenanytwothingsinanytwoconsecutiveseconds。Instrictnesstheyareidenticalandyetnotidentical,sothatinstrictnesstheyviolateafundamentalruleofstrictness——

  namely,thatathingshallneverbeitselfandnotitselfatoneandthesametime;wemustchoosebetweenlogicanddealinginapracticalspiritwithtimeandspace;itisnotsurprising,therefore,thatlogic,inspiteoftheshowofrespectoutwardlypaidtoher,istoldtostandasidewhenpeoplecometopractice。

  Inpracticeidentityisgenerallyheldtoexistwherecontinuityisonlybrokenslowlyandpiecemeal,nevertheless,thatoccasionalperiodsofevenrapidchangearenotheldtobaridentity,appearsfromthefactthatnoonedeniesthistoholdbetweenthemicroscopicallysmallimpregnateovumandthebornchildthatspringsfromit,noryet,therefore,betweentheimpregnateovumandtheoctogenarianintowhichthechildgrows;forbothovumandoctogenarianareheldpersonallyidenticalwiththenewbornbaby,andthingsthatareidenticalwiththesameareidenticalwithoneanother。

  Thefirst,then,andmostimportantelementofheredityisthatthereshouldbeunbrokencontinuity,andhencesamenessofpersonality,betweenparentsandoffspring,inneithermorenorlessthanthesamesenseasthatinwhichanyothertwopersonalitiesaresaidtobethesame。Therepetition,therefore,ofitsdevelopmentalstagesbyanyoffspringmustberegardedassomethingwhichtheembryorepeatingthemhasalreadydoneonce,inthepersonofoneorotherparent;andifonce,then,asmanytimesastherehavebeengenerationsbetweenanygivenembryonowrepeatingit,andthepointinlifefromwhichwestarted——say,forexample,theamoeba。Inthecaseofasexuallyandsexuallyproducedorganismsalike,theoffspringmustbeheldtocontinuethepersonalityoftheparentorparents,andhenceontheoccasionofeveryfreshdevelopment,toberepeatingsomethingwhichinthepersonofitsparentorparentsithasdoneonce,andifonce,thenanynumberoftimes,already。

  Itisobvious,therefore,thatthegerm-plasm(orwhateverthefancywordforitmaybe)ofanyonegenerationisasphysicallyidenticalwiththegerm-plasmofitspredecessorasanytwothingscanbe。

  ThedifferencebetweenProfessorWeismannand,wewillsay,Heringiansconsistsinthefactthatthefirstmaintainsthenewgerm-plasmwhenonthepointofrepeatingitsdevelopmentalprocessestotakepracticallynocognisanceofanythingthathashappenedtoitsincethelastoccasiononwhichitdevelopeditself;

  whilethelattermaintainthatoffspringtakesmuchthesamekindofaccountofwhathashappenedtoitinthepersonsofitsparentssincethelastoccasiononwhichitdevelopeditself,aspeopleinordinarylifetakeofthingsthathappentothem。Indailylifepeopleletfairlynormalcircumstancescomeandgowithoutmuchheedasmattersofcourse。Iftheyhavebeenluckytheymakeanoteofitandtrytorepeattheirsuccess。Iftheyhavebeenunfortunatebuthaverecoveredrapidlytheysoonforgetit;iftheyhavesufferedlonganddeeplytheygrizzleoveritandarescaredandscarredbyitforalongtime。Thequestionisoneofcognisanceornon-cognisanceonthepartofthenewgerms,ofthemoreprofoundimpressionsmadeonthemwhiletheywereonewiththeirparents,betweentheoccasionoftheirlastprecedingdevelopment,andthenewcourseonwhichtheyareabouttoenter。ThosewhoacceptthetheoryputforwardindependentlybyProfessorHeringofPrague(whoseworkonthissubjectistranslatedinmybook,\"UnconsciousMemory\"){40}andbymyselfin\"LifeandHabit,\"{41}believeincognizance,asdoLamarckiansgenerally。Weismannites,andwiththemtheorthodoxyofEnglishscience,findnon-cognisancemoreacceptable。

  IftheHeringianviewisaccepted,thatheredityisonlyamodeofmemory,andanextensionofmemoryfromonegenerationtoanother,thentherepetitionofitsdevelopmentbyanyembryothusbecomesonlytherepetitionofalessonlearnedbyrote;and,asIhaveelsewheresaid,ourviewoflifeissimplifiedbyfindingthatitisnolongeranequationof,say,ahundredunknownquantities,butofninety-nineonly,inasmuchastwooftheunknownquantitiesprovetobesubstantiallyidentical。Inthiscasetheinheritanceofacquiredcharacteristicscannotbedisputed,foritispostulatedinthetheorythateachembryotakesnoteof,remembersandisguidedbytheprofounderimpressionsmadeuponitwhileinthepersonsofitsparents,betweenitspresentandlastprecedingdevelopment。Tomaintainthisistomaintainuseanddisusetobethemainfactorsthroughoutorganicdevelopment;todenyitistodenythatuseanddisusecanhaveanyconceivableeffect。ForthedetailedreasonswhichledmetomyownconclusionsImustreferthereadertomybooks,\"LifeandHabit\"{42}and\"UnconsciousMemory,\"{42}theconclusionsofwhichhavebeenoftenadopted,butnever,thatIhaveseen,disputed。Abriefresumeoftheleadingpointsintheargumentisallthatspacewillhereallowmetogive。

  Wehaveseenthatitisafirstrequirementofhereditythatthereshallbephysicalcontinuitybetweenparentsandoffspring。Thisholdsgoodwithmemory。Theremustbecontinuedidentitybetweenthepersonrememberingandthepersontowhomthethingthatisrememberedhappened。Wecannotrememberthingsthathappenedtosomeoneelse,andinourabsence。Wecanonlyrememberhavingheardofthem。Wehaveseen,however,thatthereisasmuchbona-

  fidesamenessofpersonalitybetweenparentsandoffspringuptothetimeatwhichtheoffspringquitstheparent’sbody,asthereisbetweenthedifferentstatesoftheparenthimselfatanytwoconsecutivemoments;theoffspringtherefore,beingoneandthesamepersonwithitsprogenitorsuntilitquitsthem,canbeheldtorememberwhathappenedtothemwithin,ofcourse,thelimitationstowhichallmemoryissubject,asmuchastheprogenitorscanrememberwhathappenedearliertothemselves。Whetheritdoessoremembercanonlybesettledbyobservingwhetheritactsaslivingbeingscommonlydowhentheyareactingunderguidanceofmemory。Iwillendeavourtoshowthat,thoughheredityandhabitbasedonmemorygoaboutindifferentdresses,yetifwecatchthemseparately——fortheyareneverseentogether——andstripthemthereisnotamolenorstrawberry-mark,nortricknorleeroftheone,butwefinditintheotheralso。

  Whatarethemolesandstrawberry-marksofhabitualaction,oractionsrememberedandthusrepeated?First,themoreoftenwerepeatthemthemoreeasilyandunconsciouslywedothem。Lookatreading,writing,walking,talking,playingthepiano,&c。;thelongerwehavepractisedanyoneoftheseacquiredhabits,themoreeasily,automaticallyandunconsciously,weperformit。Look,ontheotherhand,broadly,atthethreepointstowhichIcalledattentionin\"LifeandHabit\":-

  I。Thatwearemostconsciousofandhavemostcontroloversuchhabitsasspeech,theuprightposition,theartsandsciences——whichareacquisitionspeculiartothehumanrace,alwaysacquiredafterbirth,andnotcommontoourselvesandanyancestorwhohadnotbecomeentirelyhuman。

  II。Thatwearelessconsciousofandhavelesscontrolovereatinganddrinking[providedthefoodbenormal],swallowing,breathing,seeing,andhearing——whichwereacquisitionsofourprehumanancestry,andforwhichwehadprovidedourselveswithallthenecessaryapparatusbeforewesawlight,butwhicharestill,geologicallyspeaking,recent。

  III。Thatwearemostunconsciousofandhaveleastcontroloverourdigestionandcirculation——powerspossessedevenbyourinvertebrateancestry,and,geologicallyspeaking,ofextremeantiquity。

  Ihaveputtheforegoingverybroadly,butenoughisgiventoshowthereaderthegistoftheargument。Letitbenotedthatdisturbanceanddeparture,toanyseriousextent,fromnormalpracticetendstoinduceresumptionofconsciousnesseveninthecaseofsucholdhabitsasbreathing,seeing,andhearing,digestionandthecirculationoftheblood。Soitiswithhabitualactionsingeneral。Letaplayerbeneversoproficientonanyinstrument,hewillbeputoutifthenormalconditionsunderwhichheplaysaretoowidelydepartedfrom,andwillthendoconsciously,ifindeedhecandoitatall,whathehadhithertobeendoingunconsciously。Itisanaxiomasregardsactionsacquiredafterbirth,thatweneverdothemautomaticallysaveastheresultoflongpractice;thestagesinthecaseofanyacquiredfacility,theinceptionofwhichwehavebeenabletowatch,haveinvariablybeenfromanothingnessofignorantimpotencetoalittlesomethingnessofhighlyself-

  conscious,arduousperformance,andthencetotheunselfconsciousnessofeasymastery。IsawoneyearapoorblindladofabouteighteensittingonawallbythewaysideatVarese,playingtheconcertinawithhiswholebody,andsnortinglikeachild。Thenextyeartheboynolongersnorted,andheplayedwithhisfingersonly;theyearafterthatheseemedhardlytoknowwhetherhewasplayingornot,itcamesoeasilytohim。Iknownoexceptiontothisrule。Whereistheintricateandatonetimedifficultartinwhichperfectautomaticeasehasbeenreachedexceptastheresultoflongpractice?If,then,whereverwecantracethedevelopmentofautomatismwefindittohavetakenthiscourse,isitnotmostreasonabletoinferthatithastakenthesameevenwhenithasriseninregionsthatarebeyondourken?

  Oughtwenot,wheneverweseeadifficultactionperformed,automaticallytosuspectantecedentpractice?Grantedthatwithouttheconsiderationsinregardtoidentitypresentedaboveitwouldnothavebeeneasytoseewhereababyofadayoldcouldhavehadthepracticewhichenablesittodoasmuchasitdoesunconsciously,butevenwithouttheseconsiderationsitwouldhavebeenmoreeasytosupposethatthenecessaryopportunitieshadnotbeenwanting,thanthattheeasyperformancecouldhavebeengainedwithoutpracticeandmemory。

  WhenIwrote\"LifeandHabit\"(originallypublishedin1877)Isaidinslightlydifferentwords:-

  \"Shallwesaythatababyofadayoldsucks(whichinvolvesthewholeprincipleofthepumpandhenceaprofoundpracticalknowledgeofthelawsofpneumaticsandhydrostatics),digests,oxygenisesitsblood——millionsofyearsbeforeanyonehaddiscoveredoxygen——seesandhears,operationsthatinvolveanunconsciousknowledgeofthefactsconcerningopticsandacousticscomparedwithwhichtheconsciousdiscoveriesofNewtonareinsignificant——shallwesaythatababycandoallthesethingsatonce,doingthemsowellandsoregularlywithoutbeingevenabletogivethemattention,andyetwithoutmistake,andshallwealsosayatthesametimethatithasnotlearnttodothem,andneverdidthembefore?

  \"Suchanassertionwouldcontradictthewholeexperienceofmankind。\"

  Ihavemetwithnothingduringthethirteenyearssincetheforegoingwaspublishedthathasgivenmeanyqualmsaboutitssoundness。Fromthepointofviewofthelawcourtsandeverydaylifeitis,ofcourse,nonsense;butinthekingdomofthought,asinthatofheaven,therearemanymansions,andwhatwouldbeextravaganceinthecottageorfarmhouse,asitwere,ofdailypractice,isbutcommondecencyinthepalaceofhighphilosophy,whereindwellsevolution。Ifweleaveevolutionalone,wemaysticktocommonpracticeandthelawcourts;touchevolutionandweareinanotherworld;nothigher,notlower,butdifferentasharmonyfromcounterpoint。As,however,inthemostabsolutecounterpointthereisstillharmony,andinthemostabsoluteharmonystillcounterpoint,sohighphilosophyshouldbestillintouchwithcommonsense,andcommonsensewithhighphilosophy。

  Thecommon-senseviewofthemattertopeoplewhoarenotover-

  curiousandtowhomtimeismoney,willbethatababyisnotababyuntilitisborn,andthatwhenbornitshouldbeborninwedlock。

  Nevertheless,asasoptohighphilosophy,everybabyisallowedtobetheoffspringofitsfatherandmother。

  Thehigh-philosophyviewofthematteristhateveryhumanbeingisstillbutafresheditionoftheprimordialcellwiththelatestadditionsandcorrections;therehasbeennoleapnorbreakincontinuityanywhere;themanofto-dayistheprimordialcellofmillionsofyearsagoastrulyasheisthehimselfofyesterday;hecanonlybedeniedtobetheoneongroundsthatwillprovehimnottobetheother。Everyoneisbothhimselfandallhisdirectancestorsanddescendantsaswell;therefore,ifwewouldbelogical,heisonealsowithallhiscousins,nomatterhowdistant,forheandtheyarealikeidenticalwiththeprimordialcell,andwehavealreadynoteditasanaxiomthatthingswhichareidenticalwiththesameareidenticalwithoneanother。Thisispracticallymakinghimonewithalllivingthings,whetheranimalorvegetable,thateverhaveexistedoreverwill——somethingofallwhichmayhavebeeninthemindofSophocleswhenhewrote:-

  \"NorseestthouyetthegatheringhostsofillThatshallen-onetheebothwiththineownselfAndwiththineoffspring。\"

  Andallthishascomeofadmittingthatamanmaybethesamepersonfortwodaysrunning!Asforsoppingcommonsenseitwillbeenoughtosaythattheseremarksaretobetakeninastrictlyscientificsense,andhavenoappreciableimportanceasregardslifeandconduct。Truetheydealwiththefoundationsonwhichalllifeandconductarebased,butlikeotherfoundationstheyarehiddenoutofsight,andthesoundertheyare,thelesswetroubleourselvesaboutthem。

  Whatothermaincommonfeaturesbetweenheredityandmemorymaywenotebesidesthefactthatneithercanexistwithoutthatkindofphysicalcontinuitywhichwecallpersonalidentity?First,thedevelopmentoftheembryoproceedsinanestablishedorder;somustallhabitualactionsbasedonmemory。Disturbthenormalorderandtheperformanceisarrested。Thebetterweknow\"GodsavetheQueen,\"thelesseasilycanweplayorsingitbackwards。Thereturnofmemoryagaindependsonthereturnofideasassociatedwiththeparticularthingthatisremembered——weremembernothingbutforthepresenceofthese,andwhenenoughofthesearepresentedtousweremembereverything。So,ifthedevelopmentofanembryoisduetomemory,weshouldsupposethememoryoftheimpregnateovumtorevertnottoyesterday,whenitwasinthepersonsofitsparents,buttothelastoccasiononwhichitwasanimpregnateovum。Thereturnoftheoldenvironmentandthepresenceofoldassociationswouldatonceinvolverecollectionofthecoursethatshouldbenexttaken,andthesameshouldhappenthroughoutthewholecourseofdevelopment。Theactualcourseofdevelopmentpresentspreciselythephenomenaagreeablewiththis。ForfullertreatmentofthispointImustreferthereadertothechapterontheabeyanceofmemoryinmybook\"LifeandHabit,\"alreadyreferredto。

  Secondly,werememberbestourlastfewperformancesofanygivenkind,soourpresentperformancewillprobablyresemblesomeoneorotherofthese;werememberourearlierperformancesbywayofresiduumonly,buteverynowandthenwereverttoanearlierhabit。

  Thisfeatureofmemoryismanifestedinhereditybythewayinwhichoffspringcommonlyresemblesmostitsnearerancestors,butsometimesrevertstoearlierones。Brothersandsisters,eachasitweregivingtheirownversionofthesamestory,butindifferentwords,shouldgenerallyresembleeachothermorecloselythanmoredistantrelations。Andthisiswhatactuallywefind。

  Thirdly,theintroductionofslightlynewelementsintoamethodalreadyestablishedvariesitbeneficially;thenewissoonfusedwiththeold,andthemonotonyceasestobeoppressive。Butifthenewbetooforeign,wecannotfusetheoldandthenew——natureseemingtohateequallytoowideadeviationfromordinarypracticeandnoneatall。Thisfactreappearsinheredityasthebeneficialeffectsofoccasionalcrossingontheonehand,andontheother,inthegenerallyobservedsterilityofhybrids。Ifhereditybeanaffairofmemory,howcananembryo,sayofamule,beexpectedtobuildupamuleonthestrengthofbuttwomule-memories?Hybridismcausesafaultinthechainofmemory,anditistothiscausethattheusualsterilityofhybridsmustbereferred。

  Fourthly,itrequiresmanyrepeatedimpressionstofixamethodfirmly,butwhenithasbeenengrainedintousweceasetohavemuchrecollectionofthemannerinwhichitcametobeso,orindeedofanyindividualrepetition,butsometimesasingleimpression,ifprolongedaswellasprofound,producesalastingimpressionandisliabletoreturnwithsuddenforce,andthentogoonreturningtousatintervals。Asageneralrule,however,abnormalimpressionscannotlongholdtheirownagainsttheoverwhelmingpreponderanceofnormalauthority。Thisappearsinheredityasthenormalnon-

  inheritanceofmutilationsontheonehand,andontheotherastheiroccasionalinheritanceinthecaseofinjuriesfollowedbydisease。

  Fifthly,ifheredityandmemoryareessentiallythesame,weshouldexpectthatnoanimalwoulddevelopnewstructuresofimportanceaftertheageatwhichitsspeciesbeginsordinarilytocontinueitsrace;forwecannotsupposeoffspringtorememberanythingthathappenstotheparentsubsequentlytotheparent’sceasingtocontaintheoffspringwithinitself。Fromtheaverageage,therefore,ofreproduction,offspringshouldceasetohaveanyfarthersteady,continuousmemorytofallbackupon;whatmemorythereisshouldbefulloffaults,andassuchunreliable。Anorganismoughttodevelopaslongasitisbackedbymemory——thatistosay,untiltheaverageageatwhichreproductionbegins;itshouldthencontinuetogoforatimeontheimpetusalreadyreceived,andshouldeventuallydecaythroughfailureofanymemorytosupportit,andtellitwhattodo。Thiscorrespondsabsolutelywithwhatweobserveinorganismsgenerally,andexplains,ontheonehand,whytheageofpubertymarksthebeginningofcompleteddevelopment——ariddlehithertonotonlyunexplainedbut,sofarasI

  haveseen,unasked;itexplains,ontheotherhand,thephenomenaofoldage——hithertowithoutevenattemptatexplanation。

  Sixthly,thoseorganismsthatarethelongestinreachingmaturityshouldontheaveragebethelongest-lived,fortheywillhavereceivedthemostmomentousimpulsefromtheweightofmemorybehindthem。Thisharmoniseswiththelatestopinionastothefacts。InhisarticleonWeismannintheContemporaryReviewforMay1890,Mr。

  Romaneswrites:\"ProfessorWeismannhasshownthatthereisthroughoutthemetazoaageneralcorrelationbetweenthenaturallifetimeofindividualscomposinganygivenspecies,andtheageatwhichtheyreachmaturityorfirstbecomecapableofprocreation。\"

  This,Ibelieve,hasbeentheconclusiongenerallyarrivedatbybiologistsforsomeyearspast。

  Lateness,then,intheaverageageofreproductionappearstobetheprincipleunderlyinglongevity。Theredoesnotappearatfirstsighttobemuchconnectionbetweensuchdistinctandapparentlydisconnectedphenomenaas1,theorderlynormalprogressofdevelopment;2,atavismandtheresumptionofferalcharacteristics;

  3,themoreordinaryresemblanceinterseofnearerrelatives;4,thebenefitofanoccasionalcross,andtheusualsterilityofhybrids;5,theunconsciousnesswithwhichalikebodilydevelopmentandordinaryphysiologicalfunctionsproceed,solongastheyarenormal;6,theordinarynon-inheritance,butoccasionalinheritanceofmutilations;7,thefactthatpubertyindicatestheapproachofmaturity;8,thephenomenaofmiddlelifeandoldage;9,theprincipleunderlyinglongevity。Thesephenomenahavenoconceivablebearingononeanotheruntilheredityandmemoryareregardedaspartofthesamestory。Identifythesetwothings,andIknownophenomenonofhereditythatdoesnotimmediatelybecomeinfinitelymoreintelligible。Isitconceivablethatatheorywhichharmonisessomanyfactshithertoregardedaswithouteitherconnectionorexplanationshouldnotdeserveatanyrateconsiderationfromthosewhoprofesstotakeaninterestinbiology?

  Itisnotasthoughthetheorywereunknown,orhadbeencondemnedbyourleadingmenofscience。ProfessorRayLankesterintroducedittoEnglishreadersinanappreciativenoticeofProfessorHering’saddress,whichappearedinNature,July18,1876。HewrotetotheAthenaeum,March24,1884,andclaimedcreditforhavingdoneso,butIdonotbelievehehaseversaidmoreinpublicaboutitthanwhatIhaveherereferredto。Mr。RomanesdidindeedtrytocrushitinNature,January27,1881,butin1883,inhis\"MentalEvolutioninAnimals,\"headopteditsmainconclusionwithoutacknowledgment。TheAthenaeum,tomyunboundedsurprise,calledhimtotaskforthis(March1,1884),andsincethattimehehasgiventheHeringiantheoryasufficientlywideberth。Mr。Wallaceshowedhimselffavourablyenoughdisposedtowardstheviewthatheredityandmemoryarepartofthesamestorywhenhereviewedmybook\"LifeandHabit\"inNature,March27,1879,buthehasneversincebetrayedanysignofbeingawarethatsuchatheoryexisted。Mr。

  HerbertSpencerwrotetotheAthenaeum(April5,1884),andclaimedthetheoryforhimself,but,inspiteofhisdoingthis,hehasnever,thatIhaveseen,referredtothematteragain。Ihavedealtsufficientlywithhisclaiminmybook,\"LuckorCunning。\"{43}

  Lastly,ProfessorHeringhimselfhasneverthatIknowoftouchedhisowntheorysincethesingleshortaddressreadin1870,andtranslatedbymein1881。Everyone,evenitsoriginator,exceptmyself,seemsafraidtoopenhismouthaboutit。OfcoursetheinferencesuggestsitselfthatotherpeoplehavemoresensethanI

  have。Ireadilyadmitit;butwhyhavesomanyofourleadersshownsuchastronghankeringafterthetheory,ifthereisnothinginit?

  ThedeadlockthatIhavepointedoutasexistinginDarwinismwill,Idoubtnot,leaderelongtoaconsiderationofProfessorHering’stheory。EnglishbiologistsarelittlelikelytofindWeismannsatisfactoryforlong,andifhebreaksdownthereisnothingleftforthembutLamarck,supplementedbytheimportantandelucidatorycorollaryonhistheoryproposedbyProfessorHering。Whenthetimearrivesforthistoobtainahearingitwillbeconfirmed,doubtless,byargumentsclearerandmoreforciblethananyIhavebeenabletoadduce;IshallthenbedelightedtoresignthechampionshipwhichtillthenIshallcontinue,asforsomeyearspast,tohavemuchpleasureinsustaining。Heretoforemysatisfactionhasmainlylaininthefactthatmoreofourprominentmenofsciencehaveseemedanxioustoclaimthetheorythantorefuteit;intheconfidencethusengenderedIleaveittoanyfullerconsiderationwhichtheoutlineIhaveabovegivenmayinclinethereadertobestowuponit。

  Footnotes:

  {1}PublishedintheUniversalReview,July1888。

  {2}PublishedintheUniversalReview,December1890。

  {3}PublishedintheUniversalReview,May1889。AsIhaveseveraltimesbeenaskedifthelettersherereprintedwerenotfabricatedbyButlerhimself,Itakethisopportunityofstatingthattheyareauthenticineveryparticular,andthattheoriginalsarenowinmypossession。——R。A。S。

  {4}AnaddressdeliveredattheSomervilleClub,February27,1895。

  {5}\"TheFoundationsofBelief,\"bytheRightHon。A。J。Balfour。

  Longmans,1895,p。48。

  {6}PublishedintheUniversalReview,November1888。

  {7}SincethisessaywaswrittenithasbeenascertainedbyCavaliereFrancescoNegri,ofCasaleMonferrato,thatTabachettidiedin1615。If,therefore,theSanctuaryofMontrigonewasnotfoundeduntil1631,itisplainthatTabachetticannothaveworkedthere。AllthelatestdiscoveriesaboutTabachetti’scareerwillbefoundinCavaliereNegri’spamphlet\"IlSantuariodiCrea\"

  (Alessandria,1902)。Seealsonoteonp。154。——R。A。S。

  {8}PublishedintheUniversalReview,December1889。

  {9}Longmans&Co。,1890。

  {10}Longmans&Co。,1890。

  {11}PublishedintheUniversalReview,November1890。

  {12}Longmans&Co。,1890。

  {13}M。Ruppen’swordsrun:\"1687wurdedieKapellezurhohenStiegegebaut,1747durchZusatzvergrossertund1755mitOrgelnausgestattet。AntonRuppen,eingeschickterSteinhauermidMaurermeisterleitetedenKapellebau,undmachtedarindaskleinereAltarlein。BeiderhohenStiegewarfruherkeinGebetshauslein;nureinwunderthatigesBildleinderMutterGottesstanddaineinerMauervordemfrommeHirtenundvielandachtigesVolkunterfreiemHimmelbeteten。

  \"1709wurdendiekleinenKapelleleindie15GeheimnissedesPsaltersvorstellandaufdemWegezurhohenStiegegebaut。JederHaushalterdesViertelsFeeubernahmdenBaueinesdieserGeheimnisskapellen,undeinbesondererGutthaterdieserfrommenUnternehmungwarHeinrichAndenmatten,nachherBruderderGeselischaftJesu。\"

  {14}ThestoryofTabachetti’sincarcerationisverydoubtful。

  CavaliereF。Negri,towhosebookonTabachettiandhisworkatCreaIhavealreadyreferredthereader,doesnotmentionit。TabachettilefthisnativeDinantin1585,andfromthatdateuntilhisdeathin1615heappearstohaveworkedchieflyatVaralloandCrea。

  Thereisadocumentinexistencestatingthatin1588heexecutedastatueforthehermitageofS。Rocco,atCrea,which,ifitistobereliedon,disposesbothoftheincarcerationandofthevisittoSaas。Itispossible,however,thatthedateis1598,inwhichcaseButler’stheoryofthevisittoSaasmayholdgood。In1590

  TabachettiwascertainlyatVarallo,andagainin1594,1599,and1602。Hediedin1615,possiblyduringavisittoVarallo,thoughhishomeatthattimewasCostigliole,nearAsti。——R。A。S。

  {15}ThisisthuschronicledbyM。Ruppen:\"1589den9SeptemberwareineWassergrosse,dievielSchadenverursachte。DieThalstrasse,dievondenSteinmattenanbiszurKircheamUferderVisplag,wurdeganzzerstort。ManwardgezwungeneineneueStrasseineinigerEntfernungvomWasserdurcheinenaltenFusswegauszuhauenwelchevierundeinerhalbenViertelderKlafter,oder6

  Schuhund9Zollbreitsoilte。\"(p。43)。

  {16}AlecturedeliveredattheWorkingMen’sCollegeinGreatOrmondStreet,March15,1890;rewrittenanddeliveredagainattheSomervilleClub,February13,1894。

  {17}\"CorrelationofForces\":Longmans,1874,p。15。

  {18}\"ThreeLecturesontheScienceofLanguage,\"Longmans,1889,p。4。

  {19}\"ScienceofThought,\"Longmans,1887,p。9。

  {20}PublishedintheUniversalReview,April,May,andJune1890。

  {21}\"VoyagesoftheAdventureandBeagle,\"iii。p。237。

  {22}\"Luck,orCunning,asthemainmeansofOrganicModification?\"

  (Longmans),pp。179,180。

  {23}JournalsoftheProceedingsoftheLinneanSociety(Zoology,vol。iii。),1859,p。61。

  {24}\"Darwinism\"(Macmillan,1889),p。129。

  {25}Longmans,1890,p。376。

  {26}SeeNature,March6,1890。

  {27}\"OriginofSpecies,\"sixthedition,1888,vol。i。p。168。

  {28}\"OriginofSpecies,\"sixthedition,1888,vol。ii。p。261。

  {29}Mr。J。T。Cunningham,oftheMarineBiologicalLaboratory,Plymouth,hascalledmyattentiontothefactthatIhaveascribedtoProfessorRayLankesteracriticismonMr。Wallace’sremarksupontheeyesofcertainfiat-fish,whichProfessorRayLankesterwas,inreality,onlyadopting——withfullacknowledgment——fromMr。

  Cunningham。Mr。Cunninghamhasleftittomewhethertocorrectmyomissionpubliclyornot,buthewouldsoplainlyprefermydoingsothatIconsidermyselfboundtoinsertthisnote。CuriouslyenoughIfindthatinmybook\"EvolutionOldandNew,\"IgavewhatLamarckactuallysaidupontheeyesofflat-fish,andhavingbeenledtoreturntothesubject,Imayaswellquotehiswords。Hewrote:-

  \"Need——alwaysoccasionedbythecircumstancesinwhichananimalisplaced,andfollowedbysustainedeffortsatgratification——cannotonlymodifyanorgan——thatistosay,augmentorreduceit——butcanchangeitspositionwhenthecaserequiresitsremoval。

  \"Oceanfisheshaveoccasiontoseewhatisoneithersideofthem,andhavetheireyesaccordinglyplacedoneithersideoftheirhead。

  Somefishes,however,havetheirabodenearcoastsonsubmarinebanksandinclinations,andarethusforcedtoflattenthemselvesasmuchaspossibleinordertogetasnearastheycantotheshore。

  Inthissituationtheyreceivemorelightfromabovethanfrombelow,andfinditnecessarytopayattentiontowhateverhappenstobeabovethem;thisneedhasinvolvedthedisplacementoftheireyes,whichnowtaketheremarkablepositionwhichweobserveinthecaseofsoles,turbots,plaice,&c。Thetransferofpositionisnotevenyetcompleteinthecaseofthesefishes,andtheeyesarenot,therefore,symmetricallyplaced;buttheyaresowiththeskate,whoseheadandwholebodyareequallydisposedoneithersidealongitudinalsection。Hencetheeyesofthisfishareplacedsymmetricallyupontheuppermostside。\"——PhilosophieZoologique,tom。i。,pp。250,251。EditionC。Martins。Paris,1873。

  {30}\"EssaysonHeredity,\"&c。,Oxford,1889,p。171。

  {31}\"EssaysonHeredity,\"&c。,Oxford,1889,p。266。

  {32}\"Darwinism,\"1889,p。440。

  {33}Page83。

  {34}Vol。i。p。466,&c。Ed。1885。

  {35}\"Darwinism,\"p。440。

  {36}Longmans,1890。

  {37}Tom。iv。p。383。Ed。1753。

  {38}Essays,&c。,p。447。

  {39}\"Zoonomia,\"1794,vol。i。p。480。

  {40}Longmans,1890。

  {41}Longmans,1890。

  {42}Longmans,1890。

  {43}Longmans,1890。

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