IHAVEendeavouredtoshow,intheprecedingEssay,thattheANTHROPINI,orManFamily,formaverywelldefinedgroupofthePrimates,betweenwhichandtheimmediatelyfollowingFamily,theCATARHINI,thereis,intheexistingworld,thesameentireabsenceofanytransitionalformorconnectinglink,asbetweentheCATARHINIandPLATYRHINI。
Itisacommonlyreceiveddoctrine,however,thatthestructuralintervalsbetweenthevariousexistingmodificationsoforganicbeingsmaybediminished,orevenobliterated,ifwetakeintoaccountthelongandvariedsuccessionofanimalsandplantswhichhaveprecededthosenowlivingandwhichareknowntousonlybytheirfossilizedremains。Howfarthisdoctrineiswellbased,howfar,ontheotherhand,asourknowledgeatpresentstands,itisanoverstatementoftherealfactsofthecase,andanexaggerationoftheconclusionsfairlydeduciblefromthem,arepointsofgraveimportance,butintothediscussionofwhichIdonot,atpresent,proposetoenter。Itisenoughthatsuchaviewoftherelationsofextincttolivingbeingshasbeenpropounded,toleadustoinquire,withanxiety,howfartherecentdiscoveriesofhumanremainsinafossilstatebearout,oroppose,thatview。
Ishallconfinemyself,indiscussingthisquestion,tothosefragmentaryHumanskullsfromthecavesofEngisinthevalleyoftheMeuse,inBelgium,andoftheNeanderthalnearDusseldorf,thegeologicalrelationsofwhichhavebeenexaminedwithsomuchcarebySirCharlesLyell;uponwhosehighauthorityIshalltakeitforgranted,thattheEngisskullbelongedtoacontemporaryoftheMammoth(’Elephasprimigenius’)andofthewoollyRhinoceros(’Rhinocerostichorhinus’),withthebonesofwhichitwasfoundassociated;andthattheNeanderthalskullisofgreat,thoughuncertain,antiquity。
Whateverbethegeologicalageofthelatterskull,Iconceiveitisquitesafe(ontheordinaryprinciplesofpaleontologicalreasoning)toassumethattheformertakesusto,atleast,thefurthersideofthevaguebiologicallimit,whichseparatesthepresentgeologicalepochfromthatwhichimmediatelyprecededit。AndtherecanbenodoubtthatthephysicalgeographyofEuropehaschangedwonderfully,sincethebonesofMenandMammoths,HyaenasandRhinoceroseswerewashedpell—mellintothecaveofEngis。
TheskullfromthecaveofEngiswasoriginallydiscoveredbyProfessorSchmerling,andwasdescribedbyhim,togetherwithotherhumanremainsdisinterredatthesametime,inhisvaluablework,’RecherchessurlesossemensfossilesdecouvertsdanslescavernesdelaProvincedeLiege’,publishedin1833(p。59,’etseq。’),fromwhichthefollowingparagraphsareextracted,thepreciseexpressionsoftheauthorbeing,asfaraspossible,preserved。
\"Inthefirstplace,Imustremarkthatthesehumanremains,whichareinmypossession,arecharacterizedlikethousandsofboneswhichI
havelatelybeendisinterring,bytheextentofthedecompositionwhichtheyhaveundergone,whichispreciselythesameasthatoftheextinctspecies:all,withafewexceptions,arebroken;somefewarerounded,asisfrequentlyfoundtobethecaseinfossilremainsofotherspecies。Thefracturesareverticaloroblique;noneofthemareeroded;theircolourdoesnotdifferfromthatofotherfossilbones,andvariesfromwhitishyellowtoblackish。Allarelighterthanrecentbones,withtheexceptionofthosewhichhaveacalcareousincrustation,andthecavitiesofwhicharefilledwithsuchmatter。
\"ThecraniumwhichIhavecausedtobefigured,PlateI。,Figs。1,2,isthatofanoldperson。Thesuturesarebeginningtobeeffaced:allthefacialbonesarewanting,andofthetemporalbonesonlyafragmentofthatoftherightsideispreserved。
\"Thefaceandthebaseofthecraniumhadbeendetachedbeforetheskullwasdepositedinthecave,forwewereunabletofindthoseparts,thoughthewholecavernwasregularlysearched。Thecraniumwasmetwithatadepthofametreandahalf[fivefeetnearly],hiddenunderanosseousbreccia,composedoftheremainsofsmallanimals,andcontainingonerhinocerostusk,withseveralteethofhorsesandofruminants。Thisbreccia,whichhasbeenspokenofabove(p。30),wasametre[31/4feetabout]wide,androsetotheheightofametreandahalfabovethefloorofthecavern,tothewallsofwhichitadheredstrongly。
\"Theearthwhichcontainedthishumanskullexhibitednotraceofdisturbance:teethofrhinoceros,horse,hyaena,andbear,surroundeditonallsides。
FIG。22。——TheskullfromthecaveofEngis——viewedfromtherightside。
’a’glabella,’b’occipitalprotuberance,(’a’to’b’
glabello—occipitalline),’c’auditoryforamen。
\"ThefamousBlumenbach*hasdirectedattentiontothedifferencespresentedbytheformandthedimensionsofhumancraniaofdifferentraces。Thisimportantworkwouldhaveassistedusgreatly,iftheface,apartessentialforthedeterminationofrace,withmoreorlessaccuracy,hadnotbeenwantinginourfossilcranium。
[footnote]*DecasCollectionissuaecraniorumdiversarumgentiumillustrata。Gottingae,1790—1820。
\"Weareconvincedthateveniftheskullhadbeencomplete,itwouldnothavebeenpossibletopronounce,withcertainty,uponasinglespecimen;forindividualvariationsaresonumerousinthecraniaofoneandthesamerace,thatonecannot,withoutlayingoneselfopentolargechancesoferror,drawanyinferencefromasinglefragmentofacraniumtothegeneralformoftheheadtowhichitbelonged。
\"Nevertheless,inordertoneglectnopointrespectingtheformofthisfossilskull,wemayobservethat,fromthefirst,theelongatedandnarrowformoftheforeheadattractedourattention。
\"Infact,theslightelevationofthefrontal,itsnarrowness,andtheformoftheorbit,approximateitmorenearlytothecraniumofanEthiopianthantothatofanEuropean:theelongatedformandtheproducedocciputarealsocharacterswhichwebelievetobeobservableinourfossilcranium;buttoremovealldoubtuponthatsubjectIhavecausedthecontoursofthecraniumofanEuropeanandofanEthiopiantobedrawnandtheforeheadsrepresented。PlateII。,Figs。1and2,and,inthesameplate,Figs。3and4,willrenderthedifferenceseasilydistinguishable;andasingleglanceatthefigureswillbemoreinstructivethanalongandwearisomedescription。
\"Atwhateverconclusionwemayarriveastotheoriginofthemanfromwhencethisfossilskullproceeded,wemayexpressanopinionwithoutexposingourselvestoafruitlesscontroversy。Eachmayadoptthehypothesiswhichseemstohimmostprobable:formyownpart,Iholdittobedemonstratedthatthiscraniumhasbelongedtoapersonoflimitedintellectualfaculties,andweconcludethencethatitbelongedtoamanofalowdegreeofcivilization:adeductionwhichisborneoutbycontrastingthecapacityofthefrontalwiththatoftheoccipitalregion。
\"Anothercraniumofayoungindividualwasdiscoveredinthefloorofthecavernbesidethetoothofanelephant;theskullwasentirewhenfound,butthemomentitwaslifteditfellintopieces,whichIhavenot,asyet,beenabletoputtogetheragain。ButIhaverepresentedthebonesoftheupperjaw,PlateI。,Fig。5。Thestateofthealveoliandtheteeth,showsthatthemolarshadnotyetpiercedthegum。
Detachedmilkmolarsandsomefragmentsofahumanskullproceedfromthissameplace。TheFigure3representsahumansuperiorincisortooth,thesizeofwhichistrulyremarkable。*
[footnote]*Inasubsequentpassage,Schmerlingremarksupontheoccurrenceofanincisortooth’ofenormoussize’fromthecavernsofEngihoul。Thetoothfiguredissomewhatlong,butitsdimensionsdonotappeartometobeotherwiseremarkable。
\"Figure4isafragmentofasuperiormaxillarybone,themolarteethofwhichareworndowntotheroots。
\"Ipossesstwovertebrae,afirstandlastdorsal。
\"Aclavicleoftheleftside(seePlateIII。,Fig。1);althoughitbelongedtoayoungindividual,thisboneshowsthathemusthavebeenofgreatstature。*
[footnote]*Thefigureofthisclaviclemeasures5inchesfromendtoendinastraightline——sothattheboneisratherasmallthanalargeone。
\"Twofragmentsoftheradius,badlypreserved,donotindicatethattheheightoftheman,towhomtheybelonged,exceededfivefeetandahalf。
\"Astotheremainsoftheupperextremities,thosewhichareinmypossessionconsistmerelyofafragmentofanulnaandofaradius(PlateIII。,Figs。5and6)。
\"Figure2,PlateIV。,representsametacarpalbone,containedinthebreccia,ofwhichwehavespoken;itwasfoundinthelowerpartabovethecranium:addtothissomemetacarpalbones,foundatverydifferentdistances,half—a—dozenmetatarsals,threephalangesofthehand,andoneofthefoot。
\"ThisisabriefenumerationoftheremainsofhumanbonescollectedinthecavernofEngis,whichhaspreservedforustheremainsofthreeindividuals,surroundedbythoseoftheElephant,oftheRhinoceros,andofCarnivoraofspeciesunknowninthepresentcreation。\"
FromthecaveofEngihoul,oppositethatofEngis,ontherightbankoftheMeuse,SchmerlingobtainedtheremainsofthreeotherindividualsofMan,amongwhichwereonlytwofragmentsofparietalbones,butmanybonesoftheextremities。Inonecaseabrokenfragmentofanulnawassolderedtoalikefragmentofaradiusbystalagmite,aconditionfrequentlyobservedamongthebonesoftheCaveBear(’Ursusspelaeus’),foundintheBelgiancaverns。
ItwasinthecavernofEngisthatProfessorSchmerlingfound,incrustedwithstalagmiteandjoinedtoastone,thepointedboneimplement,whichhehasfiguredinFig。7ofhisPlateXXXVI。,andworkedflintswerefoundbyhiminallthoseBelgiancaves,whichcontainedanabundanceoffossilbones。
AshortletterfromM。GeoffroySt。Hilaire,publishedinthe’ComptesRendus’oftheAcademyofSciencesofParis,forJuly2nd,1838,speaksofavisit(andapparentlyaveryhastyone)paidtothecollectionofProfessor’Schermidt’(whichispresumablyamisprintforSchmerling)
atLiege。ThewriterbrieflycriticisesthedrawingswhichillustrateSchmerling’swork,andaffirmsthatthe\"humancraniumisalittlelongerthanitisrepresented\"inSchmerling’sfigure。Theonlyotherremarkworthquotingisthis:——\"Theaspectofthehumanbonesdifferslittlefromthatofthecavebones,withwhichwearefamiliar,andofwhichthereisaconsiderablecollectioninthesameplace。Withrespecttotheirspecialforms,comparedwiththoseofthevarietiesofrecenthumancrania,few’certain’conclusionscanbeputforward;formuchgreaterdifferencesexistbetweenthedifferentspecimensofwell—characterizedvarieties,thanbetweenthefossilcraniumofLiegeandthatofoneofthosevarietiesselectedasatermofcomparison。\"
GeoffroySt。Hilaire’sremarksare,itwillbeobserved,littlebutanechoofthephilosophicdoubtsofthedescriberanddiscovereroftheremains。AstothecritiqueuponSchmerling’sfigures,Ifindthatthesideviewgivenbythelatterisreallyabout3/10thsofaninchshorterthantheoriginal,andthatthefrontviewisdiminishedtoaboutthesameextent。Otherwisetherepresentationisnot,inanyway,inaccurate,butcorrespondsverywellwiththecastwhichisinmypossession。
Apieceoftheoccipitalbone,whichSchmerlingseemstohavemissed,hassincebeenfittedontotherestofthecraniumbyanaccomplishedanatomist,Dr。Spring,ofLiege,underwhosedirectionanexcellentplastercastwasmadeforSirCharlesLyell。Itisuponandfromaduplicateofthatcastthatmyownobservationsandtheaccompanyingfigures,theoutlinesofwhicharecopiedfromveryaccurateCameralucidadrawings,bymyfriendMr。Busk,reducedtoone—halfofthenaturalsize,aremade。
AsProfessorSchmerlingobserves,thebaseoftheskullisdestroyed,andthefacialbonesareentirelyabsent;buttheroofofthecranium,consistingofthefrontal,parietal,andthegreaterpartoftheoccipitalbones,asfarasthemiddleoftheoccipitalforamen,isentireornearlyso。Thelefttemporalboneiswanting。Oftherighttemporal,thepartsintheimmediateneighbourhoodoftheauditoryforamen,themastoidprocess,andaconsiderableportionofthesquamouselementofthetemporalarewellpreserved(Fig。22)。
Thelinesoffracturewhichremainbetweenthecoadjustedpiecesoftheskull,andarefaithfullydisplayedinSchmerling’sfigure,arereadilytraceableinthecast。Thesuturesarealsodiscernible,butthecomplexdispositionoftheirserrations,showninthefigure,isnotobviousinthecast。Thoughtheridgeswhichgiveattachmenttomusclesarenotexcessivelyprominent,theyarewellmarked,andtakentogetherwiththeapparentlywelldevelopedfrontalsinuses,andtheconditionofthesutures,leavenodoubtonmymindthattheskullisthatofanadult,ifnotmiddle—agedman。
Theextremelengthoftheskullis7。7inches。Itsextremebreadth,whichcorrespondsverynearlywiththeintervalbetweentheparietalprotuberances,isnotmorethan5。4inches。Theproportionofthelengthtothebreadthisthereforeverynearlyas100to70。Ifalinebedrawnfromthepointatwhichthebrowcurvesintowardstherootofthenose,andwhichiscalledthe’glabella’(’a’)(Fig。22),totheoccipitalprotuberance(’b’),andthedistancetothehighestpointofthearchoftheskullbemeasuredperpendicularlyfromthisline,itwillbefoundtobe4。75inches。Viewedfromabove,Fig。23,A,theforeheadpresentsanevenlyroundedcurve,andpassesintothecontourofthesidesandbackoftheskull,whichdescribesatolerablyregularellipticalcurve。
Thefrontview(Fig。23,B)showsthattheroofoftheskullwasveryregularlyandelegantlyarchedinthetransversedirection,andthatthetransversediameterwasalittlelessbelowtheparietalprotuberances,thanabovethem。Theforeheadcannotbecallednarrowinrelationtotherestoftheskull,norcanitbecalledaretreatingforehead;onthecontrary,theantero—posteriorcontouroftheskulliswellarched,sothatthedistancealongthatcontour,fromthenasaldepressiontotheoccipitalprotuberance,measuresabout13。75inches。
Thetransversearcoftheskull,measuredfromoneauditoryforamentotheother,acrossthemiddleofthesagittalsuture,isabout13
inches。Thesagittalsutureitselfis5。5incheslong。
Thesupraciliaryprominencesorbrow—ridges(oneachsideof’a’,Fig。
22)arewell,butnotexcessively,developed,andareseparatedbyamediandepression。TheirprincipalelevationisdisposedsoobliquelythatIjudgethemtobeduetolargefrontalsinuses。
Ifalinejoiningtheglabellaandtheoccipitalprotuberance(’a’,’b’,Fig。22)bemadehorizontal,nopartoftheoccipitalregionprojectsmorethan1/10thofaninchbehindtheposteriorextremityofthatline,andtheupperedgeoftheauditoryforamen(’c’)isalmostincontactwithalinedrawnparallelwiththisupontheoutersurfaceoftheskull。
Atransverselinedrawnfromoneauditoryforamentotheothertraverses,asusual,theforepartoftheoccipitalforamen。Thecapacityoftheinteriorofthisfragmentaryskullhasnotbeenascertained。
ThehistoryoftheHumanremainsfromthecavernintheNeanderthalmaybestbegiveninthewordsoftheiroriginaldescriber,DrSchaaffhausen*,astranslatedbyMr。Busk。
[footnote]*ONTHECRANIAOFTHEMOSTANCIENTRACESOFMAN。
ByProfessorD。Schaaffhausen,ofBonn。(FromMuller’s’Archiv’。,1858,pp。453。)WithRemarks,andoriginalFigures,takenfromaCastoftheNeanderthalCranium。ByGeorgeBusk,F。R。S。,etc。’NaturalHistoryReview’。April,1861。
\"Intheearlypartoftheyear1857,ahumanskeletonwasdiscoveredinalimestonecaveintheNeanderthal,nearHochdal,betweenDusseldorfandElberfeld。Ofthis,however,Iwasunabletoprocuremorethanaplastercastofthecranium,takenatElberfeld,fromwhichIdrewupanaccountofitsremarkableconformation,whichwas,inthefirstinstance,readonthe4thofFebruary,1857,atthemeetingoftheLowerRhineMedicalandNaturalHistorySociety,atBonn。*
[footnote]*’Verhandl。d。Naturhist。’Vereinsderpreuss。
RheinlandeundWestphalens。,xiv。Bonn,1857。
SubsequentlyDr。Fuhlrott,towhomscienceisindebtedforthepreservationofthesebones,whichwerenotatfirstregardedashuman,andintowhosepossessiontheyafterwardscame,broughtthecraniumfromElberfeldtoBonn,andentrustedittomeformoreaccurateanatomicalexamination。AttheGeneralMeetingoftheNaturalHistorySocietyofPrussianRhinelandandWestphalia,atBonn,onthe2ndofJune,1857,*DrFuhlrotthimselfgaveafullaccountofthelocality,andofthecircumstancesunderwhichthediscoverywasmade。
[footnote]*’Ib。Correspondenzblatt。No。2。
Hewasofopinionthatthebonesmightberegardedasfossil;andincomingtothisconclusion,helaidespecialstressupontheexistenceofdendriticdeposits,withwhichtheirsurfacewascovered,andwhichwerefirstnoticeduponthembyProfessorMeyer。TothiscommunicationIappendedabriefreportontheresultsofmyanatomicalexaminationofthebones。TheconclusionsatwhichIarrivedwere:——1st。Thattheextraordinaryformoftheskullwasduetoanaturalconformationhithertonotknowntoexist,eveninthemostbarbarousraces。2nd。
ThattheseremarkablehumanremainsbelongedtoaperiodantecedenttothetimeoftheCeltsandGermans,andwereinallprobabilityderivedfromoneofthewildracesofNorth—westernEurope,spokenofbyLatinwriters;andwhichwereencounteredasautochthonesbytheGermanimmigrants。And3rdly。Thatitwasbeyonddoubtthatthesehumanrelicsweretraceabletoaperiodatwhichthelatestanimalsofthediluviumstillexisted;butthatnoproofofthisassumption,norconsequentlyoftheirso—termed’fossil’condition,wasaffordedbythecircumstancesunderwhichtheboneswerediscovered。
FIG。23。——TheEngisskullviewedfromabove(A)andinfront(B)。
\"AsDr。Fuhlrotthasnotyetpublishedhisdescriptionofthesecircumstances,Iborrowthefollowingaccountofthemfromoneofhisletters。’Asmallcaveorgrotto,highenoughtoadmitaman,andabout15feetdeepfromtheentrance,whichis7or8feetwide,existsinthesouthernwallofthegorgeoftheNeanderthal,asitistermed,atadistanceofabout100feetfromtheDussel,andabout60feetabovethebottomofthevalley。Initsearlieranduninjuredcondition,thiscavernopeneduponanarrowplateaulyinginfrontofit,andfromwhichtherockywalldescendedalmostperpendicularlyintotheriver。Itcouldbereached,thoughwithdifficulty,fromabove。
Theunevenfloorwascoveredtoathicknessof4or5feetwithadepositofmud,sparinglyintermixedwithroundedfragmentsofchert。
Intheremovingofthisdeposit,theboneswerediscovered。Theskullwasfirstnoticed,placednearesttotheentranceofthecavern;andfurtherin,theotherbones,lyinginthesamehorizontalplane。OfthisIwasassured,inthemostpositiveterms,bytwolabourerswhowereemployedtoclearoutthegrotto,andwhowerequestionedbymeonthespot。Atfirstnoideawasentertainedofthebonesbeinghuman;
anditwasnottillseveralweeksaftertheirdiscoverythattheywererecognisedassuchbyme,andplacedinsecurity。But,astheimportanceofthediscoverywasnotatthetimeperceived,thelabourerswereverycarelessinthecollecting,andsecuredchieflyonlythelargerbones;andtothiscircumstanceitmaybeattributedthatfragmentsmerelyoftheprobablyperfectskeletoncameintomypossession’
\"Myanatomicalexaminationofthesebonesaffordedthefollowingresults:——
\"Thecraniumisofunusualsize,andofalongellipticalform。Amostremarkablepeculiarityisatonceobviousintheextraordinarydevelopmentofthefrontalsinuses,owingtowhichthesuperciliaryridges,whichcoalescecompletelyinthemiddle,arerenderedsoprominent,thatthefrontalboneexhibitsaconsiderablehollowordepressionabove,orratherbehindthem,whilstadeepdepressionisalsoformedinthesituationoftherootofthenose。Theforeheadisnarrowandlow,thoughthemiddleandhinderportionsofthecranialarcharewelldeveloped。Unfortunately,thefragmentoftheskullthathasbeenpreservedconsistsonlyoftheportionsituatedabovetheroofoftheorbitsandthesuperioroccipitalridges,whicharegreatlydeveloped,andalmostconjoinedsoastoformahorizontaleminence。Itincludesalmostthewholeofthefrontalbone,bothparietals,asmallpartofthesquamousandtheupper—thirdoftheoccipital。Therecentlyfracturedsurfacesshowthattheskullwasbrokenatthetimeofitsdisinterment。Thecavityholds16,876grainsofwater,whenceitscubicalcontentsmaybeestimatedat57。64inches,or1033。24cubiccentimetres。Inmakingthisestimation,thewaterissupposedtostandonalevelwiththeorbitalplateofthefrontal,withthedeepestnotchinthesquamousmarginoftheparietal,andwiththesuperiorsemicircularridgesoftheoccipital。Estimatedindriedmillet—seed,thecontentsequalled31ounces,PrussianApothecaries’weight。Thesemicircularlineindicatingtheupperboundaryoftheattachmentofthetemporalmuscle,thoughnotverystronglymarked,ascendsneverthelesstomorethanhalftheheightoftheparietalbone。Ontherightsuperciliaryridgeisobservableanobliquefurrowordepression,indicativeofaninjuryreceivedduringlife。*
[footnote]*This,Mr。Buskhaspointedout,isprobablythenotchforthefrontalnerve。Thecoronalandsagittalsuturesareontheexteriornearlyclosed,andontheinsidesocompletelyossifiedastohaveleftnotraceswhatever,whilstthelambdoidalremainsquiteopen。ThedepressionsforthePacchionianglandsaredeepandnumerous;andthereisanunusuallydeepvasculargrooveimmediatelybehindthecoronalsuture,which,asitterminatesintheforamen,nodoubttransmitteda’venaemissaria’。Thecourseofthefrontalsutureisindicatedexternallybyaslightridge;andwhereitjoinsthecoronal,thisridgerisesintoasmallprotuberance。Thecourseofthesagittalsutureisgrooved,andabovetheangleoftheoccipitalbonetheparietalsaredepressed。
mm。*
[footnote]*ThenumbersinbracketsarethosewhichIshouldassigntothedifferentmeasures,astakenfromtheplastercast。——G。B。
Thelengthoftheskullfromthenasalprocessofthefrontaloverthevertextothesuperiorsemicircularlinesoftheoccipitalmeasures……303(300)=12。0\"。
Circumferenceovertheorbitalridgesandthesuperiorsemicircularlinesoftheoccipital……590(590)=23。37\"or23\"。
Widthofthefrontalfromthemiddleofthetemporallineononesidetothesamepointontheopposite……104(114)=4。1\"——4。5\"。
Lengthofthefrontalfromthenasal。
processtothecoronalsuture……133(125)=5。25\"——5\"。
Extremewidthofthefrontalsinuses……25(23)=1。0\"——0。9\"。
Verticalheightabovealinejoiningthedeepestnotchesinthesquamousborderoftheparietals……70=2。75\"。
Widthofhinderpartofskullfromoneparietalprotuberancetotheother……138(150)=5。4\"——5。9\"
Distancefromtheupperangleoftheoccipitaltothesuperiorsemicircularlines……51(60)=1。9\"——2。4\"。
Thicknessoftheboneattheparietalprotuberance……8。
——attheangleoftheoccipital……9。
——atthesuperiorsemicircularlineoftheoccipital……10=0。3\"
\"Besidesthecranium,thefollowingboneshavebeensecured:——
\"1。Boththigh—bones,perfect。These,liketheskull,andalltheotherbones,arecharacterizedbytheirunusualthickness,andthegreatdevelopmentofalltheelevationsanddepressionsfortheattachmentofmuscles。IntheAnatomicalMuseumatBonn,underthedesignationof’Giant’s—bones,’aresomerecentthigh—bones,withwhichinthicknesstheforegoingprettynearlycorrespond,althoughtheyareshorter。
Giant’sbones。Fossilbones。
mm。mm。
Length……542=21。4\"……438=17。4\"
Diameterofheadoffemur……54=2。14\"……53=2。0\"
\"oflowerarticularend,fromonecondyletotheother……89=3。5\"……87=3。4\"
Diameteroffemurinthemiddle……33=1。2\"……30=1。1\"
\"2。Aperfectrighthumerus,whosesizeshowsthatitbelongstothethigh—bones。
mm。
Length……312=12。3\"
Thicknessinthemiddle……26=1。0\"
Diameterofhead……49=1。9\"
\"Alsoaperfectrightradiusofcorrespondingdimensions,andtheupper—thirdofarightulnacorrespondingtothehumerusandradius。
\"3。Alefthumerusofwhichtheupper—thirdiswanting,andwhichissomuchslendererthantherightasapparentlytobelongtoadistinctindividual;aleft’ulna’,which,thoughcomplete,ispathologicallydeformed,thecoronoidprocessbeingsomuchenlargedbybonygrowth,thatflexureoftheelbowbeyondarightanglemusthavebeenimpossible;theanteriorfossaofthehumerusforthereceptionofthecoronoidprocessbeingalsofilledupwithasimilarbonygrowth。Atthesametime,theolecranoniscurvedstronglydownwards。Asthebonepresentsnosignofrachiticdegeneration,itmaybesupposedthataninjurysustainedduringlifewasthecauseoftheanchylosis。Whentheleftulnaiscomparedwiththerightradius,itmightatfirstsightbeconcludedthatthebonesrespectivelybelongedtodifferentindividuals,theulnabeingmorethanhalfaninchtooshortforarticulationwithacorrespondingradius。Butitisclearthatthisshortening,aswellastheattenuationofthelefthumerus,arebothconsequentuponthepathologicalconditionabovedescribed。
\"4。Aleft’ilium’,almostperfect,andbelongingtothefemur:afragmentoftheright’scapula’;theanteriorextremityofariboftherightside;andthesamepartofariboftheleftside;thehinderpartofariboftherightside;andlastly,twohinderportionsandonemiddleportionofribs,whichfromtheirunusuallyroundedshape,andabruptcurvature,moreresembletheribsofacarnivorousanimalthanthoseofaman。Dr。H。v。Meyer,however,towhosejudgmentIdefer,willnotventuretodeclarethemtoberibsofanyanimal;anditonlyremainstosupposethatthisabnormalconditionhasarisenfromanunusuallypowerfuldevelopmentofthethoracicmuscles。
\"Thebonesadherestronglytothetongue,although,asprovedbytheuseofhydrochloricacid,thegreaterpartofthecartilageisstillretainedinthem,whichappears,however,tohaveundergonethattransformationintogelatinewhichhasbeenobservedbyv。Bibrainfossilbones。Thesurfaceofallthebonesisinmanyspotscoveredwithminuteblackspecks,which,moreespeciallyunderalens,areseentobeformedofverydelicate’dendrites’。Thesedeposits,whichwerefirstobservedonthebonesbyDr。Meyer,aremostdistinctontheinnersurfaceofthecranialbones。Theyconsistofaferruginouscompound,and,fromtheirblackcolour,maybesupposedtocontainmanganese。Similardendriticformationsalsooccur,notunfrequently,onlaminatedrocks,andareusuallyfoundinminutefissuresandcracks。
AtthemeetingoftheLowerRhineSocietyatBonn,onthe1stApril,1857,Prof。MeyerstatedthathehadnoticedinthemuseumofPoppelsdorfsimilardendriticcrystallizationsonseveralfossilbonesofanimals,andparticularlyonthoseof’Ursusspelaeus’,butstillmoreabundantlyandbeautifullydisplayedonthefossilbonesandteethof’Equusadamiticus’,’Elephasprimigenius’,etc。,fromthecavesofBolveandSundwig。Faintindicationsofsimilar’dendrites’werevisibleinaRomanskullfromSiegburg;whilstotherancientskulls,whichhadlainforcenturiesintheearth,presentednotraceofthem。*
[footnote]*’Verh。desNaturhist’。VereinsinBonn,xiv。
1857。IamindebtedtoH。v。Meyerforthefollowingremarksonthissubject:——
’Theincipientformationofdendriticdeposits,whichwereformerlyregardedasasignofatrulyfossilcondition,isinteresting。Ithasevenbeensupposedthatindiluvialdepositsthepresenceof’dendrites’mightberegardedasaffordingacertainmarkofdistinctionbetweenbonesmixedwiththediluviumatasomewhatlaterperiodandthetruediluvialrelics,towhichaloneitwassupposedthatthesedepositswereconfined。ButIhavelongbeenconvincedthatneithercantheabsenceof’dendrites’beregardedasindicativeofrecentage,northeirpresenceassufficienttoestablishthegreatantiquityoftheobjectsuponwhichtheyoccur。Ihavemyselfnoticeduponpaper,whichcouldscarcelybemorethanayearold,dendriticdeposits,whichcouldnotbedistinguishedfromthoseonfossilbones。ThusIpossessadog’sskullfromtheRomancolonyoftheneighbouringHeddersheim,’CastrumHadrianum’,whichisinnowaydistinguishablefromthefossilbonesfromtheFrankishcaves;itpresentsthesamecolour,andadherestothetonguejustastheydo;sothatthischaracteralso,which,ataformermeetingofGermannaturalistsatBonn,gaverisetoamusingscenesbetweenBucklandandSchmerling,isnolongerofanyvalue。Indisputedcases,therefore,theconditionofthebonecanscarcelyaffordthemeansfordeterminingwithcertaintywhetheritbefossil,thatistosay,whetheritbelongtogeologicalantiquityortothehistoricalperiod。’