Andnowletusrewardourselvesforourtroublebystudyingthetragedyinthebell-glass。Clumsy,awkwardinhermovements,theScoliaslowlygoestheroundofthecircus。Atthesightofthegame,herattentionisaroused。ThestruggleisannouncedbythesamepreparationsasthosedisplayedbytheTwo-bandedScolia:theWasppolishesherwingsandtapsthetablewiththetipsofherantennae。Andview,halloo!Theattackbegins。Unabletomoveonaflatsurface,becauseofitsshortandfeeblelegs,deprivedmoreoveroftheCetonia-larva\'seccentricmeansoftravellingonitsback,theportlygrubhasnothoughtoffleeing;itcoilsitselfup。TheScolia,withherpowerfulpincers,gripsitsskinnowhere,nowelsewhere。Curvedintoacirclewiththetwoendsalmosttouching,shestrivestothrustthetipofherabdomenintothenarrowopeninginthecoilformedbythelarva。Thecontestisconductedcalmly,withoutviolentboutsateachvaryingaccident。Itisthedeterminedattemptofalivingsplitringtryingtosliponeofitsendsintoanotherlivingsplitring,whichwithequaldeterminationrefusestoopen。TheScoliaholdsthevictimsubduedwithherlegsandmandibles;shetriesoneside,thentheother,withoutmanagingtounrollthecircle,whichcontractsstillmoreasitfeelsitsdangerincreasing。Theactualcircumstancesmaketheoperationmoredifficult:thepreyslipsandrollsaboutthetablewhentheinsecthandlesittooviolently;therearenopointsofpurchaseandthestingcannotreachthedesiredspot;thefruitlesseffortsarecontinuedformorethananhour,interruptedbyperiodsofrest,duringwhichthetwoadversariesrepresenttwonarrow,interlockedrings。
WhatoughtthepowerfulCetonia-grubtodotodefytheTwo-bandedScolia,whoisfarlessvigorousthanhervictim?ItshouldimitatetheAnoxia-
larvaandremainrolleduplikeaHedgehoguntiltheenemyretires。Ittriestoescape,unrollsitselfandislost。Theotherdoesnotstirfromitspostureofdefenceandresistssuccessfully。Isthisduetoacquiredcaution?No,buttotheimpossibilityofdoingotherwiseontheslipperysurfaceofatable。Clumsy,obese,weakinthelegs,curvedintoahooklikethecommonWhiteWormThelarvaoftheCockchafer——Translator\'sNote。,theAnoxia-larvaisunabletomovealongasmoothsurface;itwritheslaboriously,lyingonitsside。Itneedstheshiftingsoilinwhich,usingitsmandiblesasaplough-share,itdigsintothegroundandburiesitself。
Letustryifsandwillshortenthestruggle,forIseenoendtoityet,aftermorethananhourofwaiting。Ilightlypowderthearena。Theattackisresumedwithavengeance。Thelarva,feelingthesand,itsnativeelement,triestoescape。Imprudentcreature!DidInotsaythatitsobstinacyinremainingrolledupwasduetonoacquiredprudencebuttothenecessityofthemoment?Thesadexperienceofpastadversitieshasnotyettaughtitthepreciousadvantagewhichitmightderivefromkeepingitscoilsclosedsolongasdangerremains。Forthatmatter,ontheunyieldingsupportofmytable,theyarenotoneandallsocautious。Thelargerseemeventohaveforgottenwhattheyknewsowellintheiryouth:thedefensiveartofcoilingthemselvesup。
Icontinuemystorywithafine-sizedspecimen,lesslikelytoslipundertheScolia\'sonslaught。Whenattacked,thelarvadoesnotcurlup,doesnotshrinkintoaringasdidthelast,whichwasyoungerandonlyhalfaslarge。Itstrugglesawkwardly,lyingonitsside,half-open。Foralldefenceittwistsabout;itopens,closesandreopensthegreathooksofitsmandibles。TheScoliagrabsitatrandom,claspsitinhershaggylegsandfornearlyaquarterofanhourbattleswiththeluscioustit-bit。Atlast,afteranotverytumultuousstruggle,whenthefavourablepositionisattainedandthepropitiousmomenthascome,thestingisimplantedinthecreature\'sthorax,inacentralpoint,belowthethroat,levelwiththefore-legs。Theeffectisinstantaneous:totalinertia,exceptoftheappendagesofthehead,theantennaeandmouth-parts。Iachievedthesameresults,thesameprickatadefinite,invariablepoint,withmyseveraloperators,renewedfromtimetotimebysomeluckycastofthenet。
Letusmention,inconclusion,thattheattackoftheInterruptedScoliaisfarlessfiercethanthatoftheTwo-bandedScolia。TheWasp,aroughsand-
digger,hasaclumsygait;hermovementsarestiffandalmostautomatic。
Shedoesnotfinditeasytorepeatherdagger-thrust。MostofthespecimenswithwhichIexperimentedrefusedasecondvictimonthefirsttwodaysaftertheirexploits。Asthoughsomnolent,theydidnotstirunlessexcitedbymyteasingthemwithabitofstraw。Althoughmoreactiveandmoreardentinthechase,theTwo-bandedScolialikewisedoesnotdrawherweaponeverytimethatIinviteher。Forallthesehuntressestherearemomentsofinactionwhichthepresenceofafreshpreyispowerlesstodisturb。
TheScoliaehavetaughtmenothingfurther,intheabsenceofsubjectsbelongingtootherspecies。Nomatter:theresultsobtainedrepresentnosmalltriumphformyideas。BeforeseeingtheScoliaeoperate,Isaid,guidedsolelybytheanatomyofthevictims,thattheCetonia-,Anoxia-andOryctes-larvaemustbeparalysedbyasinglethrustofthelancet;Ievennamedthepointwherethestingmuststrike,acentralpoint,intheimmediatevicinityofthefore-legs。Ofthethreegeneraofparalysers,twohaveallowedmetowitnesstheirsurgicalmethods,whichthethird,Ifeelcertain,willconfirm。Inbothcases,asinglethrustofthelancet;inbothcases,injectionofthevenomatapredeterminedpoint。Acalculatorinanobservatorycouldnotcomputethepositionofhisplanetwithgreateraccuracy。Anideamaybetakenasprovedwhenitattainstothismathematicalforecastofthefuture,thiscertainknowledgeoftheunknown。
Whenwilltheacclaimersofchanceachievealikesuccess?Orderappealstoorder;andchanceknowsnolaws。
CHAPTER13。THEMETHODOFTHECALICURGI。
Thenon-armouredvictims,vulnerablebythestingoveralmosttheirwholebody,ordinarycaterpillarsandLoopercaterpillars,Cetonia-andAnoxia-
larvae,whoseonlymeansofdefence,apartfromtheirmandibles,consistsofrollingsandcontortions,calledforthetestimonyofanothervictim,theSpider,almostasill-protected,butarmedwithformidablepoison-
fangs。How,inparticular,willtheRingedCalicurgussettoworkinoperatingontheBlack-belliedTarantula,theterribleLycosa,whowithasinglebitekillstheMoleortheSparrowandendangersthelifeofman?
HowdoestheboldPompilusovercomeanadversarymorepowerfulthanherself,better-equippedwithvirulentpoisonandcapableofmakingamealofherassailant?OfalltheHuntingWasps,noneriskssuchunequalconflicts,inwhichappearanceswouldproclaimtheaggressortobethevictimandthevictimtheaggressor。
Theproblemwasonedeservingpatientstudy。True,Iforesaw,fromtheSpider\'sorganization,asinglestinginthecentreofthethorax;butthatdidnotexplainthevictoryoftheWasp,emergingsafeandsoundfromhertusslewithsuchaquarry。Ihadtoseewhatoccurred。ThechiefdifficultywasthescarcityoftheCalicurgus。ItiseasyformetoobtaintheTarantulaatthedesiredmoment:thepartoftheplateauinmyneighbourhoodleftuntilledbythevine-growersprovidesmewithasmanyasarenecessary。TocapturethePompilusisanothermatter。Ihavesolittlehopeoffindingherthatspecialquestsareregardedasuseless。Tosearchforherwouldperhapsbejustthewaynottofindher。Letusrelyontheuncertaintiesofchance。ShallIgetherorshallInot?
I\'vegother。Icatchherunexpectedlyontheflowers。NextdayIsupplymyselfwithhalfadozenTarantulae。PerhapsIshallbeabletoemploythemoneaftertheotherinrepeatedduels。AsIreturnfrommyLycosa-hunt,lucksmilesuponmeagainandcrownsmydesires。AsecondCalicurgusoffersherselftomynet;sheisdraggingherheavy,paralysedSpiderbyoneleg,inthedustofthehighway。Iattachgreatvaluetomyfind:thelayingoftheegghasbecomeapressingmatter;andthemother,Ibelieve,willacceptasubstituteforhervictimwithoutmuchhesitation。Herethenaremytwocaptives,eachunderherbell-glasswithherTarantula。
Iamalleyes。Whatatragedytherewillbeinamoment!Iwait,anxiously……But……but……whatisthis?Whichofthetwoistheassailed?
Whichistheassailant?Thecharactersseemtobeinverted。TheCalicurgus,unabletoclimbupthesmoothglasswall,stridesroundtheringofthecircus。Withaproudandrapidgait,herwingsandantennaevibrating,shegoesandreturns。TheLycosaissoonseen。TheCalicurgusapproachesherwithouttheleastsignoffear,walksroundherandappearstohavetheintentionofseizingoneofherlegs。ButatthatmomenttheTarantularisesalmostverticallyonherfourhinderlegs,withherfourfrontlegsliftedandoutspread,readyforthecounterstroke。Thepoison-fangsgapewidely;adropofvenommoistenstheirtips。Theverysightofthemmakesmyfleshcreep。Inthisterribleattitude,presentingherpowerfulthoraxandtheblackvelvetofherbellytotheenemy,theSpideroverawesthePompilus,whosuddenlyturnstailandmovesaway。TheLycosathenclosesherbundleofpoisoneddaggersandresumeshernaturalpose,standingonhereightlegs;but,attheslightestattemptataggressionontheWasp\'spart,sheresumesherthreateningposition。
Shedoesmore:suddenlysheleapsandflingsherselfupontheCalicurgus;
swiftlysheclaspsherandnibblesatherwithherfangs。Withoutwieldingherstinginself-defence,theotherdisengagesherselfandmergesunscathedfromtheangryencounter。SeveraltimesinsuccessionIwitnesstheattack;andnothingseriouseverbefallstheWasp,whoswiftlywithdrawsfromthefrayandappearstohavereceivednohurt。Sheresumeshermarchingandcountermarchingnolessboldlyandswiftlythanbefore。
IsthisWaspinvulnerable,thatshethusescapesfromtheterriblefangs?
Evidentlynot。Arealbitewouldbefataltoher。Big,sturdilybuiltAcridianssuccumbLocustsandGrasshoppers——Translator\'sNote。;howisitthatshe,withherdelicateorganism,doesnot!TheSpider\'sdaggers,therefore,makenomorethananidlefeint;theirpointsdonotenterthefleshofthetight-claspedWasp。Ifthestrokeswerereal,Ishouldseebleedingwounds,Ishouldseethefangscloseforamomentonthepartseized;andwithallmyattentionIcannotdetectanythingofthekind。
ThenarethefangspowerlesstopiercetheWasp\'sinteguments?Notso。I
haveseenthempenetrate,withacracklingofbrokenarmour,thecorseletoftheAcridians,whichoffersafargreaterresistance。Onceagain,whencecomesthisstrangeimmunityoftheCalicurgusheldbetweenthelegsandassailedbythedaggersoftheTarantula?Idonotknow。Thoughinmortalperilfromtheenemyconfrontingher,theLycosathreatensherwithherfangsandcannotdecidetobite,owingtoarepugnancewhichIdonotundertaketoexplain。
Obtainingnothingmorethanalarumsandexcursionsofnogreatseriousness,Ithinkofmodifyingthegladiatorialarenaandapproximatingittonaturalconditions。Thesoilisveryimperfectlyrepresentedbymywork-table;andtheSpiderhasnotherfortress,herburrow,whichplaysapartofsomeimportancebothinattackandindefence。Ashortlengthofreedisplantedperpendicularlyinalargeearthenwarepanfilledwithsand。ThiswillbetheLycosa\'sburrow。InthemiddleIsticksomeheadsofglobe-thistlegarnishedwithhoneyasarefectoryforthePompilus;acoupleofLocusts,renewedasandwhenconsumed,willsustaintheTarantula。Thesecomfortablequarters,exposedtothesun,receivethetwocaptivesunderawire-gauzedome,whichprovidesadequateventilationforaprolongedresidence。
Myartificescometonothing;thesessioncloseswithoutresult。Adaypasses,twodays,three;stillnothinghappens。ThePompilusisassiduousinhervisitstothehoneyedflower-clusters;whenshehaseatenherfill,sheclambersupthedomeandmakesinterminablecircuitsofthenetting;
theTarantulaquietlymunchesherLocust。Iftheotherpasseswithinreach,sheswiftlyraisesherselfandwavesheroff。Theartificialburrow,thereed-stump,fulfillsitspurposeexcellently。TheLycosaandthePompilusresorttoitinturns,butwithoutquarrelling。Andthatisall。Thedramawhoseprologuewassofullofpromiseappearstobeindefinitelypostponed。
Ihavealastresource,onwhichIbasegreathopes:itistoremovemytwoCalicurgitotheverysiteoftheirinvestigationsandtoinstallthematthedooroftheSpider\'slodging,atthetopofthenaturalburrow。ItakethefieldwithanequipmentwhichIamcarryingacrossthecountryforthefirsttime:aglassbell-jar,awire-gauzecoverandthevariousimplementsneededforhandlingandtransferringmyirascibleanddangeroussubjects。
Mysearchforburrowsamongthepebblesandthetuftsofthymeandlavenderissoonsuccessful。
Hereisasplendidone。IlearnbyinsertingastrawthatitisinhabitedbyaTarantulaofasizesuitedtomyplans。Thesoilaroundtheapertureisclearedandflattenedtoreceivethewire-gauze,underwhichIplaceaPompilus。Thisisthetimetolightapipeandwait,lyingonthepebbles……Yetanotherdisappointment。Halfanhourgoesby;andtheWaspconfinesherselftotravellingroundandroundthenettingasshedidinmystudy。Shegivesnosignofgreedwhenconfrontedwiththeburrow,thoughI
canseetheTarantula\'sdiamondeyesglitteringatthebottom。
Thetrellisedwallisreplacedbytheglasswall,which,sinceitdoesnotallowhertoscaleitsheights,willobligetheWasptoremainonthegroundandatlasttotakecognizanceoftheshaft,whichsheseemstoignore。Thistimewehavedonethetrick!
Afterafewcircuitsofhercage,theCalicurgusnoticesthepityawningatherfeet。Shegoesdownit。Thisdaringconfoundsme。Ishouldneverhaveventuredtoanticipateasmuch。ThatsheshouldsuddenlyflingherselfupontheTarantulawhenthelatterisoutsideherstronghold,wellandgood;buttorushintothelair,whentheterriblemonsteriswaitingforyoubelowwiththosetwopoisoneddaggersofhers!Whatwillcomeofsuchtemerity?A
buzzingofwingsascendsfromthedepths。Runtoearthinherprivateapartments,theLycosaisnodoubtatgripswiththeintruder。ThathumofwingsistheCalicurgus\'paeanoftriumph,untilitbeherdeath-song。Theslayermaywellbetheslain。Whichofthetwowillcomeupalive?
ItistheLycosa,whohurriedlyscampersoutandpostsherselfjustovertheorificeoftheburrow,inherpostureofdefence,herfangsopen,herfourfrontlegsuplifted。Cantheotherhavebeenstabbed?Notatall,forsheemergesinherturn,notwithoutreceivingonthewayacufffromtheSpider,whoimmediatelyregainsherlair。Dislodgedfromherbasementasecondandyetathirdtime,theTarantulaalwayscomesupunwounded;shealwaysawaitsheradversaryonherthreshold,administerspunishmentandreentersherdwelling。InvaindoItrymytwoPompilialternatelyandchangetheburrow;Idonotsucceedinobservinganythingelse。Certainconditionsnotrealizedbymystratagemsarelackingtocompletethetragedy。
Discouragedbytherepetitionofmyfutileattempts,Ithrowupthegame,thericherhoweverbyonefactofsomevalue:theCalicurgus,withouttheleastfear,descendsintotheTarantula\'sdenanddislodgesher。Iimaginethatthingshappeninthesamefashionoutsidemycages。Whenexpelledfromherdwelling,theSpiderismoretimidandmorevulnerabletoattack。
Moreover,whilehamperedbyanarrowshaft,theoperatorwouldnotwieldherlancetwiththeprecisioncalledforbyherdesigns。Theboldirruptionshowsusonceagain,moreplainlythanthetusslesonmytable,theLycosa\'sreluctancetosinkherfangsintoherenemy\'sbody。Whenthetwoarefacetofaceatthebottomofthelair,thenorneveristhemomenttohaveitoutwiththefoe。TheTarantulaisinherownhouse,withallitsconveniences;everynookandcornerofthebastionisfamiliartoher。Theintruder\'smovementsarehamperedbyherignoranceofthepremises。Quick,mypoorLycosa,quick,abite;andit\'sallupwithyourpersecutor!Butyourefrain,Iknownotwhy,andyourreluctanceisthesavingoftherashinvader。ThesillySheepdoesnotreplytothebutcher\'sknifebychargingwithloweredhorns。CanitbethatyouarethePompilus\'Sheep?
Mytwosubjectsarereinstalledinmystudyundertheirwire-gauzecovers,withbedofsand,reed-stumpburrowandfreshhoney,complete。HeretheyfindagaintheirfirstLycosae,feduponLocusts。Cohabitationcontinuesforthreeweekswithoutotherincidentsthanscufflesandthreatswhichbecomelessfrequentdaybyday。Noserioushostilityisdisplayedoneitherside。AtlasttheCalicurgidie:theirdayisover。Apitifulendaftersuchanenthusiasticbeginning。
ShallIabandontheproblem?Why,notabitofit!Ihaveencounteredgreaterdifficulties,buttheyhaveneverdeterredmefromawarmly-
cherishedproject。Fortunefavoursthepersevering。Sheprovesasmuchbyofferingme,inSeptember,afortnightafterthedeathofmyTarantula-
huntresses,anotherCalicurgus,capturedforthefirsttime。ThisistheHarlequinCalicurgusC。scurra,LEP。,whosportsthesamegaudycostumeasthefirstandisalmostofthesamesize。
Nowwhatdoesthisnewcomer,ofwhomIknownothing,want?ASpider,thatiscertain;butwhich?Ahuntresslikethiswillneedacorpulentquarry:
perhapstheSilkyEpeiraE。serica,perhapstheBandedEpeiraE。
fasciata,thelargestSpidersinthedistrict,nexttotheTarantula。Thefirstofthesespreadsherlargeuprightnet,inwhichLocustsarecaught,fromoneclumpofbrushwoodtoanother。Ifindherinthecopsesontheneighbouringhills。ThesecondstretcheshersacrosstheditchesandthelittlestreamsfrequentedbytheDragon-flies。IfindherneartheAygues,besidetheirrigation-canalsfedbythetorrent。AcoupleoftripsprocuresmethetwoEpeirae,whomIoffertomycaptivenextday,bothatthesametime。Itisforhertochooseaccordingtohertaste。
Thechoiceissoonmade:theBandedEpeiraistheonepreferred。Butshedoesnotyieldwithoutprotest。OntheapproachoftheWasp,sherisesandassumesadefensiveattitude,justlikethatoftheLycosa。TheCalicurguspaysnoattentiontothreats:underherharlequin\'scoat,sheisviolentinattackandquickonherlegs。Thereisarapidexchangeoffisticuffs;andtheEpeiraliesoverturnedonherback。ThePompilusisontopofher,bellytobelly,headtohead;withherlegsshemasterstheSpider\'slegs;
withhermandiblesshegripsthecephalothorax。Shecurvesherabdomen,bringingthetipofitbeneathher;shedrawsherstingand……
Onemoment,reader,ifyouplease。Whereisthestingabouttostrike?Fromwhatwehavelearntfromtheotherparalysers,itwillbedrivenintothebreast,tosuppressthemovementofthelegs。Thatisyouropinion;itwasalsomine。Well,withoutblushingtoodeeplyatourcommonandveryexcusableerror,letusconfessthattheinsectknowsbetterthanwedo。ItknowshowtoassuresuccessbyapreparatorymanoeuvreofwhichyouandI
hadneverdreamt。Ah,whataschoolisthatoftheanimals!Isitnottruethat,beforestrikingtheadversary,youshouldtakecarenottogetwoundedyourself?TheHarlequinPompilusdoesnotdisregardthiscounselofprudence。TheEpeiracarriesbeneathherthroattwosharpdaggers,withadropofpoisonattheirpoints;theCalicurgusislostiftheSpiderbitesher。Nevertheless,heranaesthetizingdemandsperfectsteadinessofthelancet。Whatistobedoneinthefaceofthisdangerwhichmightdisconcertthemostpractisedsurgeon?Thepatientmustfirstbedisarmedandthenoperatedon。
AndinfacttheCalicurgus\'sting,aimedfrombacktofront,isdrivenintotheEpeira\'smouth,withminuteprecautionsandmarkedpersistency。Ontheinstant,thepoison-fangscloselifelesslyandtheformidablequarryispowerlesstoharm。TheWasp\'sabdomenthenextendsitsarcanddrivestheneedlebehindthefourthpairoflegs,onthemedianline,almostatthejunctionofthebellyandthecephalothorax。Atthispointtheskinisfinerandmoreeasilypenetrablethanelsewhere。Theremainderofthethoracicsurfaceiscoveredwithatoughbreast-platewhichthestingwouldperhapsfailtoperforate。Thenerve-centres,thesourceoftheleg-
movements,aresituatedalittleabovethewoundedpoint,buttheback-to-
frontdirectionofthestingmakesitpossibletoreachthem。Thislastwoundresultsintheparalysisofalltheeightlegsatonce。
Toenlargeuponitfurtherwoulddetractfromtheeloquenceofthisperformance。Firstofall,tosafeguardtheoperator,astabinthemouth,thatpointsoterriblyarmed,themostformidableofall;then,tosafeguardthelarva,asecondstabinthenerve-centresofthethorax,tosuppressthepowerofmovement。IcertainlysuspectedthattheslayersofrobustSpiderswereendowedwithspecialtalents;butIwasfarfromexpectingtheirboldlogic,whichdisarmsbeforeitparalyses。SotheTarantula-huntressmustbehave,who,undermybell-glasses,refusedtosurrenderhersecret。Inowknowwhathermethodis;ithasbeendivulgedbyacolleague。ShethrowstheterribleLycosauponherback,pricksherprickersbystingingherinthemouthandthen,incomfort,withasinglethrustofthelancet,obtainsparalysisofthelegs。
IexaminetheEpeiraimmediatelyaftertheoperationandtheTarantulawhentheCalicurgusisdraggingherbyonelegtoherburrow,atthefootofsomewall。Foralittlewhilelonger,aminuteatmost,theEpeiraconvulsivelymovesherlegs。Solongasthesethroescontinue,thePompilusdoesnotreleaseherprey。Sheseemstowatchtheprogressoftheparalysis。WiththetipsofhermandiblessheexplorestheSpider\'smouthseveraltimesover,asthoughtoascertainifthepoison-fangsarereallyinnocuous。Whenallmovementsubsides,thePompilusmakesreadytodragherpreyelsewhere。ItisthenItakechargeofit。
Whatstrikesmemorethananythingelseistheabsoluteinertiaofthefangs,whichIticklewithastrawwithoutsucceedinginrousingthemfromtheirtorpor。Thepalpi,ontheotherhand,theirimmediateneighbours,waveattheleasttouch。TheEpeiraisplacedinsafety,inaflask,andundergoesafreshexaminationaweeklater。Irritabilityhasinpartreturned。Underthestimulusofastraw,Iseeherlegsmovealittle,especiallythelowerjoints,thetibiaeandtarsi。Thepalpiareevenmoreirritableandmobile。Thesedifferentmovements,however,arelackinginvigourandcoordination;andtheSpidercannotemploythemtoturnover,muchlesstoescape。Asforthepoison-fangs,Istimulatetheminvain:I
cannotgetthemtoopenoreventostir。Theyarethereforeprofoundlyparalysedandinaspecialmanner。Thepeculiarinsistenceofthestingwhenthemouthwasstabbedtoldmeasmuchinthebeginning。
AttheendofSeptember,almostamonthaftertheoperation,theEpeiraisinthesamecondition,neitherdeadnoralive:thepalpistillquiverwhentouchedwithastraw,butnothingelsemoves。Atlength,aftersixorsevenweeks\'lethargy,realdeathsupervenes,togetherwithitscomrade,putrefaction。
TheTarantulaoftheRingedCalicurgus,asItakeherfromtheowneratthemomentoftransportation,presentsthesamepeculiarities。Thepoison-fangsarenolongerirritablewhentickledwithmystraw:afreshproof,addedtothoseofanalogy,toshowthattheLycosa,liketheEpeira,hasbeenstunginthemouth。Thepalpi,ontheotherhand,areandwillbeforweekshighlyirritableandmobile。Iwishtoemphasisethispoint,theimportanceofwhichwillberecognizedpresently。
IfounditimpossibletoprovokeasecondattackfrommyHarlequinCalicurgus:thetediumofcaptivitydidnotfavourtheexerciseofhertalents。Moreover,theEpeirasometimeshadsomethingtodowithherrefusals;acertainrusedeguerrewhichwastwiceemployedbeforemyeyesmaywellhavebaffledtheaggressor。Letmedescribetheincident,ifonlytoincreaseourrespectalittleforthesefoolishSpiders,whoareprovidedwithperfectedweaponsanddonotdaretomakeuseofthemagainsttheweakerbutbolderassailant。
TheEpeiraoccupiesthewallofthewire-gauzecage,withhereightlegswide-spreaduponthetrelliswork;theCalicurgusiswheelingroundthetopofthedome。Seizedwithpanicatthesightoftheapproachingenemy,theSpiderdropstotheground,withherbellyupwardsandherlegsgatheredtogether。Theotherdashesforward,claspsherroundthebody,exploresherandpreparestostingherinthemouth。Butshedoesnotbareherweapon。I
seeherbendingattentivelyoverthepoisonedfangs,asthoughtoinvestigatetheirterriblemechanism;shethengoesaway。TheSpiderisstillmotionless,somuchsothatIreallybelieveherdead,paralysedunknowntome,atamomentwhenIwasnotlooking。Itakeherfromthecagetoexaminehercomfortably。Nosoonerissheplacedonthetablethanbehold,shecomestolifeagainandpromptlyscampersoff!ThecunningcreaturewasshammingdeathbeneaththeWasp\'sstiletto,soartfullythatI
wastakenin。Shedeceivedanenemymorecunningthanmyself,thePompilus,whoinspectedherverycloselyandtookherforacorpseunworthyofherdagger。Perhapsthesimplecreature,liketheBearinthefableofold,alreadynoticedthesmellofhighmeat。
Thisruse,ifruseitbe,appearstomemoreoftenthannottoturntothedisadvantageoftheSpider,whetherTarantula,Epeiraoranother。TheCalicurguswhohasjustputtheSpideronherbackafterabriskfightknowsquitewellthatherprostratefoeisnotdead。Thelatter,thinkingtoprotectitself,simulatestheinertiaofacorpse;theassailantprofitsbythistodeliverhermostperilousblow,thestabinthemouth。Werethefangs,eachtippedwithitsdropofpoison,toopenthen;weretheytosnap,togiveadesperatebite,thePompiluswouldnotdaretoexposethetipofherabdomentotheirdeadlyscratch。Theshammingofdeathisexactlywhatenablesthehuntresstosucceedinherdangerousoperation。
Theysay,OguilelessEpeirae,thatthestruggleforlifehastaughtyoutoadoptthisinertattitudeforpurposesofdefence。Well,thestruggleforlifewasaverybadcounsellor。Trustrathertocommonsenseandlearn,bydegrees,atyourowncost,thattohitback,aboveallifyoucandosopromptly,isstillthebestwaytointimidatetheenemy。Fabredoesnotbelieveintheactualshammingofdeathbyanimals。Cf。“TheGlow-wormandOtherBeetles,“byJ。HenriFabre,translatedbyAlexanderTeixeiradeMattos:chapters8to15——Translator\'sNote。
Theremainderofmyobservationsontheseinsectsunderglassislittlemorethanalongseriesoffailures。OftwooperatorsonWeevils,one,theSandyCercerisC。arenaria,persistentlyscornedthevictimsoffered;theother,Ferrero\'sCercerisC。Ferreri,allowedherselftobeemptedaftertwodays\'captivity。Hertacticalmethod,asIexpected,ispreciselythatoftheCleonus-huntress,theGreatCerceris,withwhommyinvestigationscommenced。WhenconfrontedwiththeAcorn-weevil,sheseizestheinsectbythesnout,whichisimmenselylongandshapedlikeapipe-stem,andplantsherstinginitsbodytotherearoftheprothorax,betweenthefirstandsecondpairoflegs。Itisneedlesstoinsist:thespoileroftheCleonihastaughtusenoughaboutthismodeofoperationanditsresults。
NoneoftheBembex-wasps,whetherchosenamongthehuntressesoftheGadflyoramongtheloversoftheHouse-flyrabble,satisfiedmyaspirations。
TheirmethodisasunknowntomenowasatthedistantperiodwhenIusedtowatchitintheBoisdesIssards。Cf。“TheHuntingWasps“:chapters14
to18——Translator\'sNote。Theirimpetuousflight,theirloveoflongjourneysareincompatiblewithcaptivity。Stunnedbycollidingwiththewallsoftheirglassorwire-gauzeprison,theyallperishwithintwenty-
fourhours。Swifterintheirmovementsandapparentlysatisfiedwiththeirhoneyedthistle-heads,theSpheges,huntressesofCricketsorEphippigers,dieasquicklyofnostalgia。AllIofferthemleavesthemindifferent。
NorcanIgetanythingoutoftheEumenes,notablythebiggestofthem,thebuilderofgravelcupolas,Amedeus\'Eumenes。AllthePompili,excepttheHarlequinCalicurgus,refusemySpiders。ThePalarus,whopreysuponanindefinitenumberoftheHymenopteronclan,refusestotellmeifshedrinksthehoneyoftheBees,asdoesthePhilanthus,orifsheletstheothersgowithoutmanipulatingthemtomakethemdisgorge。TheTachytesdonotvouchsafetheirLocustsaglance;Stizusruficornispromptlygivesuptheghost,disdainingthePrayingMantiswhichIprovideforher。
Whatistheuseofcontinuingthislistofchecks?Therulemaybegatheredfromthesefewexamples:occasionalsuccessesandmanyfailures。Whatcanbethereason?WiththeexceptionofthePhilanthus,temptedfromtimetotimebyabumperofhoney,thepredatoryWaspsdonothuntontheirownaccount;theyhavetheirvictualling-time,whentheegg-layingisimminent,whenthefamilycallsforfood。Outsidetheseperiods,thefinestheadsofgamemightwellleavethesenectar-bibbersindifferent。Iamcarefultherefore,asfaraspossible,tocapturemysubjectsattheproperseason;
Igivepreferencetomotherscaughtuponthethresholdoftheburrowwiththeirpreybetweentheirlegs。Thisdiligenceofminebynomeansalwayssucceeds。Therearedemoralizedinsectswhich,onceunderglass,evenafterabriefdelay,nolongercareabouttheequivalentoftheirprize。
Allthespeciesdonotperhapspursuetheirgamewiththesameardour;moodandtemperamentaremorevariableeventhanconformation。Tothesefactors,whichareofthenicestorder,wemayaddthatofthehour,whichisoftenunfavourablewhenthesubjectiscaughtathaphazardontheflowers,andweshallhavemorethanenoughtoexplainthefrequencyofthefailures。Afterall,Imustbewareofrepresentingmyfailuresastherule:whatdoesnotsucceedonedaymayverywellsucceedanotherday,underdifferentconditions。Withperseveranceandalittleskill,anyonewhocarestocontinuetheseinterestingstudieswill,Iamsure,fillupmanygaps。Theproblemisdifficultbutnotimpossible。
Iwillnotquitmybell-jarswithoutsayingawordontheentomologicaltactofthecaptiveswhentheydecidetoattack。Oneofthepluckiestofmysubjects,theHairyAmmophila,wasnotalwaysprovidedwiththehereditarydishofherfamily,theGreyWorm。Iofferedherindiscriminatelyanybare-
skinnedcaterpillarsthatIchancedtofind。Somewereyellow,somegreen,somebrownwithwhiteedges。Allwereacceptedwithouthesitation,providedthattheywereofsuitablesize。Tastygamewasrecognizedwonderfullyunderverydissimilarliveries。ButayoungZeuzera-caterpillar,dugoutofthebranchesofalilac-tree,andasilkwormofsmalldimensionsweredefinitelyrefused。Theover-fedproductsofoursilkworm-nurseriesandthemystery-lovingcaterpillarwhichgnawstheinnerwoodofthelilacinspiredherwithsuspicionanddisgust,despitetheirbareskin,whichfavouredthesting,andtheirshape,whichwassimilartothatofthevictimsaccepted。
Anotherardenthuntress,theInterruptedScolia,refusedtheCetonia-grub,whichisoflikehabitswiththeAnoxia-larva;theTwo-bandedScoliaalsorefusedtheAnoxia。ThePhilanthus,theheadlongmurderessofBees,sawthroughmytrickerywhenIconfrontedherwiththeVirgilianBee,theEristalisE。tenax。She,aPhilanthus,takethisFlyforaBee!Whatnext!Thepopularideaismistaken;antiquitytooismistaken,aswitnessthe“Georgics,“whichmaketheputridremainsofasacrificedBullgivebirthtoaswarm;buttheWaspmakesnomistake。Inhereyes,whichseefartherthanours,theEristalisisanodiousDipteron,aloverofcorruption,andnothingmore。
CHAPTER14。OBJECTIONSANDREJOINDERS。
Noideaofanyscopecanbeginitssoaringflightbutstraightwaythecurmudgeonsareafterit,eagertobreakitswingsandtostampthewoundedthingunderfoot。MydiscoveryofthesurgicalmethodsthatgivetheHuntingWaspstheirpreservedfoodstuffshasundergonethecommonrule。Lettheoriesbediscussed,byallmeans:therealmoftheimaginationisanuntilleddomain,inwhicheveryoneisfreetoplanthisownconceptions。
Butrealitiesarenotopentodiscussion。Itisabadpolicytodenyfactswithnomoreauthoritythanone\'swishtofindthemuntrue。NoonethatI
knowofhasimpugnedbycontraryobservationswhatIhavesolongbeensayingabouttheanatomicalinstinctoftheWaspsthathunttheirprey;
instead,Iammetwitharguments。Mercyonus!Firstuseyoureyesandthenyoushallhaveleavetoargue!And,topersuadepeopletousetheireyes,I
meantoreply,sincewehavetimetospare,totheobjectionswhichhavebeenormayberaised。Ofcourse,Ipassoverinsilencethoseinwhichchildishdisparagementshowsitsnosetooplainly。
Thesting,Iamtold,isdirectedatonepointratherthananotherbecausethatistheonlyvulnerablepoint。Theinsectcannotchoosewhatwounditwillinflict;itstingswhereitmust。Itswonderfuloperativemethodisthenecessaryresultofthevictim\'sstructure。Letusfirst,ifweattachanyimportancetolucidity,cometoanunderstandingabouttheword“vulnerable。“Doyoumeanbythisthatthepointorratherpointswoundedbythestingaretheonlypointsatwhichalesionwillsuddenlycauseeitherdeathorparalysis?Ifso,Ishareyouropinion;notonlydoIshareit,butIwasthefirsttoproclaimit。Mywholethesisiscontainedinthat。Yes,ahundredtimesyes,thepointswoundedaretheonlyvulnerablepoints;theyareevenveryvulnerable;theyaretheonlypointswhichlendthemselvestotheinflictionofsuddendeathorelseparalysis,accordingtotheoperator\'sintention。
Butthisisnothowyouunderstandthematter:youmeanaccessibletothesting,inaword,penetrable。Herewepartcompany。Ihaveagainstme,I
admit,theWeevilsandtheBuprestesoftheCerceres。Thesemailedoneshardlygivethestingachance,savebehindtheprothorax,thepointatwhichthelancetisactuallydirected。IfIwereonetostandontrifles,I
mightobservethatinfrontoftheprothorax,underthethroat,isanaccessiblespotandthattheCercereswillhavenothingtodowithit。Butletusproceed;Igiveupthehorn-cladBeetle。
WhatarewetosayoftheGreyWormandothercaterpillarsbelovedoftheAmmophilae?Herearevictimsaccessibletothestingunderneath,ontheback,onthesides,foreandaft,everywherewiththesamefacility,exceptingthetopofthehead。Andofthisinfinityofpoints,whichareequallypenetrable,theWaspselectsten,alwaysthesame,differinginnowayfromtherest,unlessitbebythecloseproximityofthenerve-
centres。WhatarewetosayoftheCetonia-andAnoxia-larvae,whicharealwaysattackedinthefirstthoracicsegment,afterlongandpainfulstruggles,whentheassailantcanstingthegrubfreelyatwhateverpointshechooses,sinceitisquitenakedandoffersnogreaterresistancetothelancetatonepointthanatanother?
WhatarewetothinkoftheSphex\'CricketsandEphippigers,stabbedthreetimesonthesideofthethorax,whichisfairlywelldefended,whereastheabdomen,softandbulky,intowhichthestingwouldsinklikeaneedleintoapatofbutter,isneglected?DonotletusforgetthePhilanthus,whotakesnoaccounteitherofthefissuresbeneaththeabdominalplatesorofthewidehiatusbehindthecorselet,butplungesherweapon,atthebaseofthethroat,throughagapofafractionofamillimetre。LetusjustmentiontheMantis-huntingTachytes。Doesshemakeforthemostundefendedpointwhenshestabs,firstofall,atitsbase,theMantis\'dreadfulengine——thearm-pieceseachfittedwithadoublesaw——attheriskofbeingseized,transfixedandcrunchedonthespotifshemissesherblow?Whydoesshenotstrikeatthecreature\'slongabdomen?Thatwouldbequiteeasyandfreefromdanger。
AndtheCalicurgi,ifyouplease。Aretheyalsounskilledduelists,plungingthedirkintotheonlyeasilyaccessiblepoint,whentheirveryfirstmoveistoparalysethepoison-fangs?IfthereisonepointabouttheTarantulaandtheEpeirathatisdangerousanddifficulttoattack,itiscertainlythemouthwhichbiteswithitstwopoisonedharpoons。Andthesedesperadoesdaretobravethatdeadlytrap!Whydotheynotfollowyourjudiciousadvice?Theyshouldstingtheplumpbelly,whichiswhollyunprotected。Theydonot;andtheyhavetheirreasons,ashavetheothers。
All,fromthefirsttothelast,showus,clearaswaterfromtherock,thattheouterstructureofthevictimsoperatedoncountsfornothinginthemethodofoperating。Thisisdeterminedbytheinneranatomy。Thepointswoundedarenotstungbecausetheyaretheonlypointspenetrablebythelancet;theyarestungbecausetheyfulfilanimportantcondition,withoutwhichpenetrabilitylosesitsvalue。Thisconditionisnoneotherthantheimmediateproximityofthenerve-centreswhoseinfluencehastobesuppressed。Whenatclosequarterswithherprey,whethersoftorarmour-
clad,thehuntressbehavesasifsheunderstoodthenervoussystembetterthananyofus。Thethoughtlessobjectionabouttheonlypenetrablepointsis,Ihope,sweptasideforever。
Iamalsotold:
“Itispossible,ifitcomestothat,forthestingtobedeliveredintheneighbourhoodofthenerve-centres;inavictimatmostthreeorfourcentimetreslong,distancesareverysmall。Butacasualthereorthereaboutsisaverydifferentthingfromtheprecisionofwhichyouspeak。“
Oh,theyare“thereabouts,“arethey?Weshallsee!Youwantfigures,millimetres,fractions?Youshallhavethem!
FirstIcalltowitnesstheInterruptedScolia。Ifthereadernolongerhashermethodofoperatinginmind,Iwillbeghimtorefreshhismemory。Thetwoadversaries,inthepreliminaryconflict,maybefairlywellrepresentedbytworingsinterlockednotinthesameplanebutatrightangles。TheScoliagripsapointoftheAnoxia-grub\'sthorax;shecurvesherbodyunderneathitand,whileencirclingthegrub,gropeswiththetipofherabdomenalongthemedianlineofthelarva\'sneck。Owingtohertransversalposition,theassailantisnowfreetoaimherweaponinaslightlyslantingdirection,whethertowardstheheadortowardsthethorax,atthesamepointofentryinthelarva\'sthroat。Betweenthetwooppositeslantsofthesting,whichisitselfveryshort,whatcanthedistancebe?Twomillimetres。078inch——Translator\'sNote。,perhapsless。Thatisverylittle。Nomatter:lettheoperatormakeamistakeofthislength——negligible,youmaytellme——letthestingslanttowardstheheadinsteadofslantingtowardsthethorax;andtheresultoftheoperationwillbeentirelydifferent。Withaslanttowardsthehead,thecerebralgangliaarewoundedandtheirlesioncausessuddendeath。ThisisthestrokeofthePhilanthus,whokillsherBeebystingingherfrombelow,underthechin。TheScolianeededamotionlessbutnotdeadvictim,onethatwouldsupplyfreshvictuals;shewillnowhaveonlyacorpse,whichwillsoongobadandpoisonthelarva。
Withaslanttowardsthethorax,thestingwoundsthelittlemassofnerve-
cellsinthethorax。Thisistheregulationstroke,theonewhichwillinduceparalysisandleavethesmallamountoflifeneededtokeeptheprovisionsfresh。Amillimetrehigherkills;amillimetrelowerparalyses。
OnthistinydeviationthesalvationoftheScoliaracedepends。Youneednotfearthattheoperatorwillmakeanymistakeinthismicrometricalperformance:herstingalwaysslantstowardsthethorax,althoughtheoppositeinclinationisjustaspracticableandeasy。Whatwouldbetheoutcomeofathereorthereaboutsundertheseconditions?Veryoftenacorpse,aformoffoodfataltothegrub。
TheTwo-bandedScoliastingsalittlelowerdown,onthelineofdemarcationbetweenthefirsttwothoracicsegments。HerpositionislikewisetransversalinrelationtotheCetonia-grub;butthedistanceofthecervicalgangliafromthepointwherethestingenterswouldpossiblynotallowtheweaponturnedtowardstheheadtoinflictalesionfollowedbysuddendeathasintheaboveinstance。Iamcallingthiswitnesswithanotherobject。Itisextremelyunusualfortheoperator,nomatterwhatherpreyorhermethod,tomakeaslightmistakeandstingmerelysomewhereneartherequisitepoint。Iseethemallgropingwiththetipoftheabdomen,sometimesseekingpersistently,beforeunsheathing。Theythrustonlywhenthepointbeneaththestingispreciselythatatwhichthewoundwillproduceitsfulleffect。TheTwo-bandedScoliainparticularwillstrugglewiththeCetonia-grubforhalfanhouratatimetoenableherselftodriveinthestilettoattherightspot。
Weariedbyanendlessscuffle,oneofmycaptivescommittedbeforemyeyesaslightblunder,anunprecedentedthing。Herweaponenteredalittletooneside,notquiteamillimetrefromthecentralpointandstill,ofcourse,onthelineofdemarcationbetweenthefirsttwothoracicsegments。
Iatoncelaidholdofthepreciousspecimen,whichwastoteachmecuriousmattersabouttheeffectsofanill-deliveredstroke。IfImyselfhadmadetheinsectstingatthisorthatpoint,therewouldhavebeennoparticularinterestinit:theScolia,heldbetweenthefinger-tips,wouldwoundatrandom,likeaBeedefendingherself;herundirectedstingwouldinjectthepoisonathaphazard。Buthereeverythinghappenedbyrule,exceptforthelittleerrorofposition。
Well,thevictimofthisclumsyoperationhasitslegsparalysedonlyontheleftside,thesidetowardswhichtheweaponwasdeflected;itisacaseofhemiplegia。Thelegsontherightsidemove。Iftheoperationhadbeenperformedinthenormalfashiontheresultwouldhavebeensuddeninertiaofallsixlegs。Thehemiplegia,itistruedoesnotlastlong。Thetorporofthelefthalfrapidlygainstherighthalfofthebodyandthecreatureliesmotionless,incapableofburyingitselfinthemould,without,however,realizingtheconditionsindispensabletothesafetyoftheeggortheyounggrub。IfIseizeoneofitslegsorapointoftheskinwiththetweezers,itsuddenlyshrivelsandcurlsupandswellsoutagain,asitdoeswhenincompletepossessionofitsenergies。Whatwouldbecomeofanegglaidonsuchvictuals?Atthefirstclosingofthisruthlessvice,atthefirstcontraction,itwouldbecrushed,oratleastdetachedfromitsplace;andanyeggremovedfromthepointwherethemotherhasfasteneditisboundtoperish。Itneeds,ontheCetonia\'sabdomen,ayieldingsupportwhichthebitesofthenew-bornlarvawillnotsetaquiver。Theslightlyeccentricstinggivesnoneofthissoftmassoffat,alwaysoutstretchedandquiescent。Onlyonthefollowingday,afterthetorporhasmadeprogress,doesthelarvabecomesuitablyinertandlimp。Butitistoolate;andinthemeantimetheeggwouldbeinseriousdangeronthishalf-paralysedvictim。Thesting,bystrayinglessthanamillimetre,wouldleavetheScoliawithoutprogeny。
Ipromisedfractions。Heretheyare。LetusconsidertheTarantulaandtheEpeiraonwhomtheCalicurgihavejustoperated。Thefirstthrustofthestingisdeliveredinthemouth。Inbothvictimsthepoison-fangsareabsolutelylifeless:ticklingwithabitofstrawneveroncesucceedsinmakingthemopen。Ontheotherhand,thepalpi,theirverynearneighbours,theiradjunctsasitwere,possesstheircustomarymobility。Withoutanyprevioustouches,theykeeponmovingforweeks。Inenteringthemouththestingdidnotreachthecervicalganglia,orsuddendeathwouldhaveensuedandweshouldhavebeforeoureyescorpseswhichwouldgobadinafewdays,insteadoffreshcarcasesinwhichtracesofliferemainmanifestforalongtime。Thecephalicnerve-centreshavebeenspared。
Whatiswoundedthen,toprocurethisprofoundinertiaofthepoison-fangs?
Iregretthatmyanatomicalknowledgeleavesmeundecidedonthispoint。
Arethefangsactuatedbyaspecialganglion?Aretheyactuatedbyfibresissuingfromcentresexercisingfurtherfunctions?IleavetoanatomistsequippedwithmoredelicateinstrumentsthanIthetaskofelucidatingthisobscurequestion。Thesecondconjectureappearstomethemoreprobable,becauseofthepalpi,whosenerves,itseemstome,musthavethesameoriginasthoseofthefangs。Basingourargumentonthislatterhypothesis,weseethattheCalicurgushasonlyonemeansofsuppressingthemovementofthepoisonedpincerswithoutaffectingthemobilityofthepalpi,aboveallwithoutinjuringthecephaliccentresandthusproducingdeath,namely,toreachwithherstingthetwofibresactuatingthefangs,fibresasfineasahair。
Iinsistuponthispoint。Despitetheirextremedelicacy,thesetwofilamentsmustbeinjureddirectly;for,ifitwereenoughforthestingtoinjectitspoison“thereorthereabouts,“thenervesofthepalpi,soclosetothefirst,wouldundergothesameintoxicationastheadjacentregionandwouldleavethoseappendagesmotionless。Thepalpimove;theyretaintheirmobilityforaconsiderableperiod;theactionofthepoison,therefore,isevidentlysituatedinthenervesofthefangs。Therearetwoofthesenerve-filaments,veryfine,verydifficulttodiscover,evenbytheprofessionalanatomist。TheCalicurgushastoreachthemoneaftertheother,tomoistenthemwithherpoison,possiblytotransfixthem,inanycasetooperateupontheminaveryrestrictedmanner;sothatthediffusionofthevirusmaynotinvolvetheadjoiningparts。Theextremedelicacyofthissurgeryexplainswhytheweaponremainsinthemouthsolong;thepointofthestingisseekingandeventuallyfindsthetinyfractionofamillimetrewherethepoisonistoact。Thisiswhatwelearnfromthemovementsofthepalpiclosetothemotionlessfangs;theytellusthattheCalicurgiarevivisectorsofalarmingaccuracy。
Ifweacceptthehypothesisofaspecialnerve-centreforthemandibles,thedifficultywouldbealittleless,withoutdetractingfromtheoperator\'stalent。Thestingwouldthenhavetoreachabarelyvisiblespeck,anatominwhichweshouldhardlyfindroomforthepointofaneedle。Thisisthedifficultywhichthevariousparalyserssolveinordinarypractice。Dotheyactuallywoundwiththeirdirkstheganglionwhoseinfluenceistobedoneawaywith?Itispossible,butIhavetriednotesttomakesure,theinfinitelytinywoundappearingtobetoodifficulttodetectwiththeopticalinstrumentsatmydisposal。Dotheyconfinethemselvestolodgingtheirdropofpoisonontheganglion,oratalleventsinitsimmediateneighbourhood?Idonotsayno。
Ideclaremoreover,that,toprovokelightningparalysis,thepoison,ifitisnotdepositedinsidethemassofnervoussubstance,mustactfromsomewhereverynear。ThisassertionismerelyechoingwhattheTwo-bandedScoliahasjustshownus:herCetonia-grub,stunglessthanamillimetrefromtheregularspot,didnotbecomemotionlessuntilnextday。Thereisnodoubt,judgingbythisinstance,thattheeffectofthevirusspreadsinalldirectionswithinaradiusofsomeextent;butthisdiffusionisnotenoughfortheoperator,whorequiresforheregg,whichissoontobelaid,absolutesafetyfromtheveryfirst。
Ontheotherhand,theactionsoftheparalysersargueaprecisesearchfortheganglia,atalleventsforthefirstthoracicganglion,themostimportantofall。TheHairyAmmophila,amongothers,affordsusanexcellentexampleofthismethod。Herthreethrustsinthecaterpillar\'sthoraxandespeciallythelast,betweenthefirstandsecondpairoflegs,aremoreprolongedthanthestabsdistributedamongtheabdominalganglia。
Everythingjustifiesusinbelievingthat,forthesedecisiveinoculations,thestingseeksoutthecorrespondingganglionandactsonlywhenitfindsitunderitspoint。Ontheabdomenthispeculiarinsistenceceases;thestingpassesswiftlyfromonesegmenttoanother。Forthesesegments,whicharelessdangerous,theAmmophilaperhapsreliesonthediffusionofhervenom;inanycase,theinjections,thoughhastilyadministered,donotdivergefromaclosevicinityoftheganglia,fortheirfieldofactionisverylimited,asisprovedbythenumberofinoculationsnecessarytoinducecompletetorpor,or,moresimply,bythefollowingexample。
AGreyWormwhichhadjustreceiveditsfirststingonthethirdthoracicsegmentrepulsestheAmmophilaandwithajerkhurlshertoadistance。I
profitbytheoccasionandtakeholdofthegrub。Thelegsofthisthirdsegmentonlyareparalysed;theothersretaintheirusualmobility。Howeverhelplessinthetwoinjuredlegs,theanimalcanwalkverywell;itburiesitselfintheearth,returningtothesurfaceatnighttognawthestumpoflettucewithwhichIhaveservedit。Forafortnightmyparalyticretainsperfectlibertyofaction,exceptinthesegmentoperatedon;thenitdies,notofitswoundbutaccidentally。Allthistimetheeffectofthepoisonhasnotspreadbeyondtheinoculatedsegment。
Atanypointwherethestingenters,anatomyinformsusofthepresenceofanervousnucleus。Isthiscentredirectlysmittenbytheweapon?Orisitpoisonedwithvirus,fromaverysmalldistance,bytheprogressiveimpregnationoftheneighbouringtissues?Thisisthedoubtfulpoint,thoughitdoesnotinanywayinvalidatetheprecisionoftheabdominalinjections,whicharecomparativelyneglected。Asforthoseinthecaterpillar\'sthorax,theirprecisionisbeyonddispute。AftertheAmmophilae,theScoliaeand,aboveall,theCalicurgi,isitreallynecessarytobringintocourtyetotherwitnesses,whowouldallswearthat,withmodificationsofdetail,themovementoftheirlancetisstrictlyregulatedbythenervoussystemoftheprey?Thisoughttobeenough。Theproofisestablishedforthosewhohaveearstohearwith。
Othersdelightinobjectionswhoseodditysurprisesme。TheyseeinthepoisonoftheHuntingWaspsanantisepticliquidandinvictualsstoredintheirburrowspreservedmeatswhicharekeptfreshnotbyaremnantoflifebutbythevirusanditsmicrobes。Come,mylearnedmasters,letusjusttalkthematterover,betweenourselves。HaveyoueverseenthelarderofaskilledHuntingWasp,aSphexforinstance,aScolia,anAmmophila?Youhaven\'t,haveyou?Ithoughtasmuch。Yetitwouldbebettertobeginbydoingso,beforebringingthepreservativemicrobeonthescene。Theslightestexaminationwouldhaveshownyouthatthevictualscannotbecomparedexactlywithsmokedhams。Thethingmoves,thereforeitisnotdead。Thereyouhavethewholematter,initsartlesssimplicity。Thepalpimove,themandiblesopenandshut,thetarsiquiver,theantennaeandtheabdominalfilamentswavetoandfro,theabdomenthrobs,theintestinerejectsitscontents,theanimalreactstothestimulusofaneedle,allofwhichsignsarehardlycompatiblewiththeideaofpickledmeat。
HaveyouhadthecuriositytolookthroughthepagesinwhichIsetforththedetailedresultsofmyobservations?Youhaven\'t,haveyou?Again,I
thoughtasmuch。Itisapity。Youwouldtherefind,inparticular,thehistoryofcertainEphippigerswho,afterbeingstungbytheSphexaccordingtorule,wererearedbymyselfbyhand。Youmustagreethatthesearequeerpreservestobeproducedbytheuseofanantisepticfluid。TheyacceptthemouthfulswhichIofferthemonthetipofastraw;theyfeed,theysitupandtakenourishment。Ishallneverlivetoseetinnedsardinesdoingasmuch。
Iwillavoidtediousrepetitionandcontentmyselfwithaddingtomyoldsheafofproofsafewfactswhichhavenotyetbeenrelated。TheNest-
buildingOdynerusshowedusinhercellsafewChrysomela-larvaefixedbythehinderparttothesideofthereed。Thegrubfastensitselfinthiswaytothepoplar-leaftoobtainapurchasewhenthemomenthascomeforleavingthelarvalslough。Donotthesepreparationsforthenymphosistellusplainlythatthecreatureisnotdead?
TheHairyAmmophilaaffordsusanevenbetterexample。Anumberofcaterpillarsoperatedonbeforemyeyesattained,somesooner,somelater,thechrysalisstage。Mynotesareexplicitonthesubjectofsomeofthem,takenonVerbascumsinuatum。Sacrificedonthe14thofApril,theywerestillirritablewhentickledwithastrawafortnightafter。Alittlelater,thepale-greencolouringoftheearlystagesisreplacedbyareddishbrown,exceptontwoorthreesegmentsofthemedianventralsurface。Theskinwrinklesandsplits,butdoesnotcomedetachedofitsownaccord。Icaneasilyremoveitinshreds。Underthissloughappearsthefirm,chestnut-brownhornintegumentofthechrysalis。ThedevelopmentofthenymphosisissocorrectthatforamomentthecrazyhopeoccurstomethatImayseeaTurnip-mothcomeoutofthismummy,thevictimofadozendagger-thrusts。Fortherest,thereisnoattemptatspinningacocoon,nojetofsilkythreadsflungoutbythecaterpillarbeforeturningintoachrysalis。Perhapsundernormalconditionsmetamorphosistakesplacewithoutthisprotection。However,themothwhomIexpectedtoseewasbeyondthelimitsofthepossible。InthemiddleofMay,amonthaftertheoperationonthecaterpillars,mythreechrysalids,stillincompleteunderneath,inthethreeorfourmiddlesegments,witheredandatlastwentmouldy。Istheevidenceconclusivethistime?Whocanconceivesuchasillyideaasthatapreyreallydead,acorpsepreservedfromputrefactionbyanantiseptic,couldcontainwhatisperhapsthemostdelicateworkoflife,thedevelopmentofthegrubintotheperfectinsect?
Thetruthmustbedrivenintorecalcitrantbrainswithgreatblowsofthesledge-hammer。Letusoncemoreemploythismethod。InSeptemberIunearthfromaheapofmouldfiveCetonia-grubs,paralysedbytheTwo-bandedScoliaandbearingontheabdomentheasyetunhatchedeggoftheWasp。Iremovetheeggsandinstallthehelplesscreaturesonabedofleaf-mouldwithaglasscover。IproposetoseehowlongIcankeepthemfresh,abletomovetheirmandiblesandpalpi。AlreadythevictimsofvariousHuntingWaspshadinstructedmeonasimilarmatter;Iknewthattracesoflifelingerfortwo,three,fourweeksandlonger。Forinstance,IhadseentheEphippigersoftheLanguedocianSphexcontinuethewavingoftheirantennaeandtheirparalyticshuddersforfortydaysofartificialfeedingbyhand;andIusedtowonderwhetherthemoreorlessearlydeathoftheothervictimswasnotduetolackofnourishmentquiteasmuchastotheoperationwhichtheyhadundergone。However,theinsectinitsadultformusuallyhasaverybriefexistence。Itsoondies,killedbythemerefactofliving,withoutanyotheraccident。Alarvaispreferablefortheseinvestigations。Itsconstitutionislivelier,betterabletosupportprotractedabstinence,aboveallduringthewintertorpor。TheCetonia-grub,aregularlumpofbacon,nourishedbyitsownfatduringthewinterseason,fulfilstheneedfulconditionstoperfection。Whatwillbecomeofit,lyingbellyupwardsonitsbedofleaf-mould?Willitsurvivethewinter?
Attheendofamonth,threeofmygrubsturnbrownandlapseintorottenness。Theothertwokeepperfectlyfreshandmovetheirantennaeandpalpiatthetouchofastraw。Thecoldweathercomesandticklingnolongerelicitsthesesignsoflife。Theinertiaiscomplete;neverthelesstheirappearanceremainsexcellent,withoutatraceofthebrownishtinge,thesignofdeterioration。Atthereturnofthewarmweather,inthemiddleofMay,thereisasortofresurrection。Ifindmytwolarvaeturnedover,bellydownwards;muchmore:theyarehalf-buriedinthemould。Whenteased,theycoiluplazily;theymovetheirlegsaswellastheirmouth-parts,butslowlyandwithoutvigour。Thentheirstrengthseemstorevive。Theconvalescent,resuscitatedgrubsdigwithclumsyeffortsintotheirbedofmould;theydiveintoitanddisappeartoadepthofabouttwoinches。
Recoveryseemstobeimminent。
Iammistaken。InJuneIunearththeinvalids。Thistime,thelarvaearedead;theirbrowncolourtellsmeasmuch。Iexpectedbetterthings。Nevermind:thisisnotriflingsuccess。Forninemonths,ninelongmonths,thegrubsstabbedbytheScoliakeptfreshandalive。Towardstheend,torporwasdispelled,strengthandmovementreturned,sufficientlytoenablethemtoleavethesurfacewhereIhadplacedthemandtoregainthedepthsbyboringapassagethroughthesoil。Ireallythinkthatafterthisresurrectiontherewillbenomoretalkofantiseptics,unlessanduntiltinnedHerringsbegintofrolicintheirbrine。Thesubjectofthisandtheprecedingchaptersiscontinuedinanessayentitled“ThePoisonoftheBee“forwhichcf。“Bramble-beesandOthers“:chapter11——Translator\'sNote。
INDEX。
Acorn-weevil。
Amedeus\'Eumenes。
AmelesdecolorseeGreyMantis。
Ammophilaseealsothevarietiesbelow。
AmmophilahursutaseeHairyAmmophila。
AmmophilaholosericaseeSilkyAmmophila。
AmmophilaJuliiseeJules\'Ammophila。
AmmophilasabulosaseeSandyAmmophila。
AnathemaTachytes。
Anoxiaseealsothevarietiesbelow。
Anoxiaaustralis。
AnoxiamatutinalisseeMorningAnoxia。
AnoxiavillosaseeShaggyAnoxia。
Ant。
Anthidiumseealsothevarietiesbelow。
Anthidiumbellicosum。
Anthidiumscapulare。
Anthidiumseptemdentatum。
Anthophora。
AnthraxseealsoAnthraxsinuata。
Anthraxsinuata。
Ape。
AphisseePlant-louse。
Ass。
Astata。
BalaninusseealsoBalaninusglandum。
BalaninusglandumseeAcorn-weevil。
BandedEpeira。
Bat。
BeeseealsoBumble-bee,Hive-bee,Mason-bee。
Bee-eatingPhilanthus。
Beetle。
Bembexseealsothevarietiesbelow。
BembexbidentataseeTwo-prongedBembex。
BembexrostrataseeRostrateBembex。
Black,AdamandCharles。
Black-belliedTarantula。
BlackSpiderseeCellarSpider。
BlackTachytes。
Blister-beetleseeOil-beetle。
Bluebottle。
BlueOsmia。
Bombylius。
Boyle,Robert。
Brachycera。
BrachyderespubescensseePubescentBrachyderes。
Breguet,Louis。
Brillat-Savarin,Anthelme。
Brown-wingedSolenius。
Bug。
Bull。
Bull,theauthor\'sDog。
Bullock。
Bumble-bee。
Buprestis。
Buprestis-huntingCerceris。
BurntZonitis。
Butterfly。
CabbagePieris。
CalicurgusseePompilusandthevarietiesbelow。
CalicurgusannulatusseeRingedCalicurgus。
CalicurgusscurraseeHarlequinCalicurgus。
Callot,Jacques。
Cantharides。
Carpenter-bee。
CellarSpider。
Centuryco。
CercerisseealsoBuprestis-huntingCercerisandthevarietiesbelow。
CercerisarenariaseeSandCerceris。
CercerisFerreriseeFerrero\'sCerceris。
CercerisornataseeOrnateCerceris。
CerceristuberculataseeGreatCerceris。
Cerocoma。
Cetoniaseealsothevarietiesbelow。
CetoniaaurataseeGoldenCetonia。
Cetoniamorio。
Chaffinch。
ChalicodomaseeMason-bee。
Chaoucho-grapaouseeNightjar。
Chimpanzee。
ChrysomelapopuliseePoplarLeaf-beetle。
Cicada。
Cicadella。
CleonusseealsoCleonusophthalmicus。
Cleonusophthalmicus。
Cneorhinus。
Cockchafer。
ColpainterruptaseeInterruptedScolia。
CommonCockchaferseeCockchafer。
CommonWasp。
Cotton-beeseeAnthidiumscapulare。
Cow。
Crab。
CrabroseeHornet。
CrabrochrysostomusseeGolden-mouthedHornet。
Cricket。
CrownedPhilanthus。
Cuckoo。
Darwin,CharlesRobert。
Davidthepainter。
David,FelicienCesar。
Death\'s-headHawk-moth。
DevilkinseeEmpusa。
Dicranuravinula。
DioxyscinctaseeGirdledDioxys。
DogseealsoBull。
Drone-fly。
Dufour,JeanMarieLeon。
Duges,LouisAntoine。
Earth-worm。
Eight-spottedPompilus。
Empusa。
Epeiraseealsothevarietiesbelow。
EpeirafasciataseeBandedEpeira。
EpeirasericaseeSilkyEpeira。
Ephippiger。
EristalisE。tenaxseeDrone-fly。
Eucera。
EuchloraJulii。
EumenesseealsoAmedeusEumenes。
Fabricius,JohanChristian。
Favier,theauthor\'sfactotum。
Ferrero\'sCerceris。
Field-mouse。
FlyseealsoGad-fly,House-fly。
Fox。
Frog。
Gad-fly。
Galileo。
GardenScolia。
GardenSpiderseeEpeira。
Geonomus。
GirdledDioxys。
Gnat。
Goat。
GoatsuckerseeNightjar。
GoldenCetonia。
Golden-crestedWren。
Golden-mouthedHornet。
GoldenOsmia。
Gorilla。
Grasshopper。
GreatCellarSpiderseeCellarSpider。
GreatCerceris。
GreyMantis。
GreyWorm。
HairyAmmophila。
Halictus。
HarlequinCalicurgus。
Hedgehog。
Helophiluspendulus。
HemorrhoidalScolia。
Hen。
Herring。
Hive-bee。
Hog。
HornetseealsoGolden-mouthedHornet。
House-fly。
InterruptedScolia。
Jules,Ammophila。
Klug。
Lalande,JosephJeromeLeFrancaisde。
Lamellicorn。
LanguedocianSphex。
Lark。
Latreille,PierreAndre。
Leucopsisgigas,L。grandis。
Lily-beetle。
Linnet。
Locust。
Looper。
LycosaseeBlack-belliedTarantula。
MacmillanCo。
MantisseealsoGreyMantis,PrayingMantis。
Mantis-huntingTachytesseeMantis-killingTachytes。
Mantis-killingTachytes。
Mariotte,Edme。
Mason-beeseealsotheAnthophoraandthevarietiesbelow。
Mason-beeofthePebblesseeMason-beeoftheWalls。
Mason-beeoftheSheds。
Mason-beeoftheShrubs。
Mason-beeoftheWalls。
Measuring-wormseeLooper。
Megachilesericans。
Melanophora。
MeloeseeOil-beetle。
Miall,Bernard。
Midge。
MithradatesVI。
Mole。
Mole-cricket。
Monkey。
MonocerosseeOryctesnasicornis。
MorningAnoxia。
Mosquito。
Moth。
Mule。
MuscidseeHouse-fly。
Mylabris。
NarbonneLycosaseeBlack-belliedTarantula。
Nest-buildingOdynerus。
Nightjar。
Nut-weevil。
OdynerusseealsoNest-buildingOdynerus。
Oil-beetle。
OrnateCerceris。
Oryctesnasicornis。
OryctesSilenus。
Osmiaseealsothevarietiesbelow。
OsmiacyaneaseeBlueOsmia。
Osmiacyanoxantha。
OsmiaLatreilliiseeLatreille\'sOsmia。
OsmiaparvulaseeTinyOsmia。
OsmiatricornisseeThree-hornedOsmia。
Ostrich。
Otiorhynchus。
PalarusseealsoPalarusflavipes。
Palarusflavipes。
Pangonia。
Panzer\'sTachytes。
Paragus。
Pascal,Blaise。
Passerini。
Pea-weevil。
Pelopaeus。
Pentodonpunctatus。
Perez,J。
Phaneropteronfalcata。
Philanthusseealsothevarietiesbelow。
PhilanthusapivorusseeBee-eatingPhilanthus。
PhilanthuscoronatusseeCrownedPhilanthus。
PhilanthusraptorseeRobberPhilanthus。
Phynotomus。
PierisseeCabbagePieris。
Pig。
Pine-chafer。
Pithecanthropus。
Plant-louse。
Pompilusseealsothevarietiesbelow。
PompilusannulatusseeRingedCalicurgus。
Pompilusapicalis。
PompilusoctopunctatusseeEight-spottedPompilus。
PoplarLeaf-Beetle。
PrayingMantis。
PubescentBrachyderes。
Rat。
Resin-beeseeAnthidiumbellicosum,Anthidiumseptemdentatum。
RhinocerosBeetleseeOryctesnasicornis。
Rhynchitesbetuleti。
RingedCalicurgus。
RingedPompilusseeRingedCalicurgus。
RobberPhilanthus。
Robber-fly。
Robin。
Romanes,GeorgeJohn。
Rose-chaferseeCetonia,GoldenCetonia。
RostrateBembex。
SandCerceris。
SandyAmmophila。
SapygapunctataseeSpottedSapyga。
Sarcophaga。
Scarabaeid。
Scarabaeuspentodon。
Scoliaseealsothevarietiesbelow。
ScoliabifasciataseeTwo-bandedScolia。
ScoliahaemorrhoidalisseeHemorrhoidalScolia。
ScoliahortorumseeGardenScolia。
ScoliainterruptaseeInterruptedScolia。
Screech-owl。
Seal。
SegestriaperfidiaseeCellarSpider。
ShaggyAnoxia。
Sheep。
Silkworm。
SilkyAmmophila。
SilkyEpeira。
SilkyLeaf-cutterseeMegachilesericans。
Sitones。
Skua。
Slug。
Snail。
Socrates。
SoleniusfascipennisseeBrown-wingedSolenius。
SoleniusvagusseeWanderingSolenius。
Sparrow。
Sparrow-hawk。
Sphaerophoria。
SphexseealsoLanguedocianSphex,White-bandedSphex,Yellow-wingedSphex。
SpiderseealsoBlack-belliedTarantula,CellarSpider,Epeira。
SpottedSapyga。
SpurgeHawk-moth。
Stizusseealsothevarietiesbelow。
Stizusruficornis。
Stizustridentatus。
Strophosomus。
Swallow。
Swammerdam,Jan。
Syrittaperpens。
Syrphus。
TachytesseealsoMantis-killingTachytesandthevarietiesbelow。
TachytesanathemaseeAnathemaTachytes。
TachytesnigraseeBlackTachytes。
TachytesPanzeriseePanzer\'sTachytes。
TachytestarsinaseeTarsalTachytes。
Tachytesunicolor。
TarantulaseeBlack-belliedTarantula。
TarsalBembex。
TarsalTachytes。
TeixeiradeMattos,Alexander。
Three-hornedOsmia。
TinyOsmia。
Toad。
Toricelli,Evangelista。
Toussenel,Alphonse。
Turkey。
TurnipMoth。
Two-bandedScolia。
Two-prongedBembex。
Unwin,T。Fisher,Ltd。
VespacrabroseeHornet。
VirgilianBee,Virgil\'sBeeseeDrone-fly。
WanderingSolenius。
WaspseeCommonWasp。
WeevilseealsoAcorn-weevil,Nut-weevil,Pea-weevil。
Whale。
WhippoorwillseeNightjar。
White-bandedSphex。
WhiteWorm。
Wolf。
Yellow-wingedSphex。
Zeuzera。
ZonitispraeustaseeBurntZonitis。
End